Final report coastal energy impact program contract number 80-63 : a continuation of the biological inventory of the coastal zone in Lucas, Wood, Ottawa, Sandusky, Erie Lake, and Ashtabula Counties to determine locations of significant plant species, plant communities and natural areas which are threatened due to new or expanding energy facilities or activities

[From the U.S. Government Printing Office, www.gpo.gov]

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FINAL REPORT

COASTAL ENERGY IMPACT PROGRAM

Contract Number 80-63

A Continuation of the Biological Inventory of the Coastal
Zone in Lucas, Wood, Ottawa, Sandusky, Erie, Lake,
and Ashtabula Counties to Determine Locations of
Significant Plant Species, Plant Communities and
Natural Areas which are Threatened Due to New
or Expanding Energy Facilities or Activities

LU. S. DEPARTMENT OF COMMERCE NOAA
COAS1 AL SERVICES CENTER
Submitted to: "234 SUIH HOBSON AVENUE
ChAhLESTON , SC 29405-2413

Ohio Department of Energy
Coastal Energy Impact Program
30 East Broad Street, 34th Floor
Columbus, Ohio 43215

COASTAl. ZONE

INFORMATION CENTER

By:

Ohio Department of Natural Resources
Division of Natural Areas and Preserves
Fountain Square, Building F
Columbus, Ohio 43224

TD February 13, 1981
195
.E5
F562
1981

PROJECT STAFF

Ohio Department of Natural Resources
Division of Natural Areas and Preserves

Dennis M. Anderson - Research Coordinator
Barbara K. Andreas - Field Botanist, C.E.I.P.
Nneka Atiba - Business Coordinator
David M. Brandenburg - Field Botanist, C.E.I.P.
Allison W. Cusick - Staff Botanist
Anne R. Filbert - Data Processing Specialist
Patricia D. Jones - Data Systems Supervisor
Susan E. Laverty - Secretary
Robert M. McCance, Jr. - Project Administrator
Richard E. Moseley, Jr. - Division Chief
J. Steven Shelly - Data Processing Intern

AC KNOWLEDGMENTS

The Ohio Department of Natural Resources, Division of Natural Areas and

Preserves, wishes to thank the staff of the Ohio Department of Energy for the

assistance they provided in the implementation of this project. In addition,

this project would not have been possible without the support and cooperation

of the Ohio Coastal Zone Management Section. Special thanks are due to the

many landowners who granted permission for Division staff to survey their

property.

This document was funded in Part throucih the
Coastal Zone Management Act of 1972, as amend-
ed (Coastal Energy Impact Program), provided
by the U. S. Dept. of Commerce, NOAA, through
the Ohio Department of Energy.

TABLE OF CONTENTS

Page Number

PROJECT STAFF i

ACKNOWLEDGMENTS ii

TABLE OF CONTENTS iii

INTRODUCT ION 1

DATA COLLECTION SYSTEM 3

SURVEY RESULTS 5

FEDERALLY LISTED SPECIES 8

MANAGEMENT CONSIDERATIONS 9

PROTECTION STRATEGY 10

SUMMARY AND CONCLUSIONS 12

APPENDICES

A. PROJECT AUTHORIZATION LETTER 13

B. RARE SPECIES OF NATIVE OHIO WILD PLANTS 15

C. COUNTY LISTS OF RARE PLANTS OF THE COASTAL
ZONE 37

D. OHIO PLANTS OF FEDERAL CONCERN 43

E. ADMINISTRATIVE RULES FOR O.R.C. CHAPTER 1518 45

F. DATA SERVICES BROCHURE AND APPLICATION FORM 47

INTRODUCT ION

The Coastal Energy Impact Program (CEIP), created by a 1976 amendment to

Section 308 of the U.S. Coastal Zone Management Act of 1972, is a federally

funded program designed to assist coastal states and units of local government

in mitigation of on-shore effects of new or expanding coastal energy activity.

The Ohio Department of Energy has responsibility for development, implementa-

tion, and administration of Section 308 CEIP within the State of Ohio.

On July 1, 1979, the Ohio CEIP provided a Planning Grant of $22,294 in

federal funds to the Division of Natural Areas and Preserves (DNAP). Planning

grants are awarded on an 80% federal/20% non-federal funding basis. These

grants may be used to defray costs of study and planning for economic, environ-

mental, or social impacts occurring or likely to occur in the coastal zone as

a result of the siting, construction, expansion, or operation of new (since

July 26, 1976) or expanded energy facilities.

On June 12, 1980, the Ohio CEIP provided written authorization to proceed

with a second CEIP Planning Grant, Number 80-63, "A Continuation of the

Biological Inventory of the Coastal Zone in Lucas, Wood, Ottawa, Sandusky,

Erie, Lake and Ashtabula Counties to Determine Locations of Significant Plant

Species, Plant Communities and Natural Areas which are Threatened Due to New

or Expanding Energy Facilities or Activities." This grant of $12,910 was

intended to be the spring and early summer equivalent of the 1979 grant, which

covered the field season beginning on August 6, 1979. As originally proposed,

the field survey portion of the 1980 project would have been initiated in

mid-April. Due to difficulties encountered in acquiring formal federal

approval, the Division was not authorized to initiate the project until June 12,

1980. Field surveys were conducted until mid-September.

Reviewers of this report should refer to the final report for CEIP

Contract Number 79-16, dated December 28, 1979, for additional information

relating to the overall coastal zone study.

This final report is intended to supplement the 1979 project report and

to provide information which was unavailable for the 1979 report. This

report does provide a listing of state official endangered and threatened

plant species known to occur in the Ohio coastal zone. However, this report

does not provide details about rare plant occurrences, relative quality or

location of the most significant plant communities or natural areas, or

Division strategy for future preservation efforts of specific sites. Accu-

rate, detailed information can be provided to legitimate data users through

the Natural Heritage Program data services system. This system will provide

the data user with the most current information available.

DATA COLLECTION SYSTEM

Data on endangered species and significant natural plant communities in

the Lake Erie coastal zone were obtained from all major available sources to

the level of detail allowed by funding and time limitations of the 1979 contract.

Data sources included herbarium specimens, literature records, knowledgeable

individuals, and original field work. These data were organized by the data

processing systems of the Natural Heritage Program within the Ohio Department

of Natural Resources, Division of Natural Areas and Preserves.

Most of the preexisting endangered species data pertaining to the Lake

Erie coastal zone had been obtained by the Natural Heritage Program prior to

work on the 1979 CEIP contract. These data were refined, updated and reorganized

to assist field survey efforts on the project and to make the data more useful

to the Division. Attempts then were made to validate these records in the

field during 1979 and 1980.

Most of the new or improved data obtained by the Division of Natural Areas

and Preserves was a result of the efforts of the field botanists. Mr. David

Brandenburg worked part-time on the western basin (Lucas to Erie counties) from

May 12 (prior to the approved starting date of the project) until August 24.

Dr. Barbara K. Andreas worked part-time on the eastern basin from May 12 (prior

to the approved starting date of the project) until September 19. These

people were assisted by Mr. Allison Cusick, staff botanist for the Natural

Heritage Program. The pre-contract field work was conducted at the expense

of the Division and that data also is included in this report.

Confronted with the formidable task of surveying as much of the coastal

zone as possible, the field botanists concentrated on areas where previous

records, literature and personal leads, probable habitats, and intuition

directed them. Their primary goal was the location of endangered plant

species, with possibly significant plant community stands being recorded when

encountered. Specimens of state endangered, threatened, and potentially

threatened species were obtained when populations were large enough to

warrant collection. Specimens were deposited primarily at the herbarium of

The Ohio State University, with duplicates, when collected, usually being

deposited at Kent State University and/or Miami University. Sight records

(versus collection records) were made for very small populations and for

populations of species already represented by adequate collections in given

areas.

All endangered species occurrence data were organized into the Natural

Heritage Program data processing system. Each record was checked for complete-

ness and accuracy of information, and then mapped and entered into computer

storage. Such storage allows easy access to, alteration of, and addition to

the data contained on the Lake Erie coastal zone. This data system made

it possible to generate the species lists attached as appendices to this

report.

SURVEY RESULTS

The Lake Erie coastal zone contains numerous locations of state endangered

and threatened plants, as well as many significant plant communities and nat-

ural areas. Previous data collection efforts in the coastal zone have been

extensive when compared to efforts in other parts of Ohio, yet there were a

number of areas which required additional surveys. This project provided an

opportunity for the Division of Natural Areas and Preserves to verify older

data and to improve upon vague or inaccurate information about the early

summer flora of the coastal zone.

Many previously known locations of rare plants were surveyed and verified

as a result of this project. Older records were updated, improved and recon-

firmed. New locations for rare plants were discovered and better population

data was obtained on known sites. It is not possible to give exact statistics

on the number of new rare plant occurrences identified as a result of the

1980 CEIP project because it is not always possible to determine whether an

old, vague record is being updated and improved, or whether a previously

unreported occurrence has been discovered.

Research efforts in the coastal zone as a result of the 1979 CEIP con-

tract resulted in 161 new and/or improved records of rare plant occurrences

being added to the Natural Heritage Program data base. The 1980 CEIP contract

research, as well as data collected by Division staff from early May until

June 12, 1980, provided approximately 50 records of rare plant occurrences

being added to the Natural Heritage Program data base. Of particular

significance was the discovery of one and possibly two presumed extirpated

plant species in the coastal zone during 1980.

A list of the rare plants of Ohio is attached as Appendix B of this

report. Of the 565 Ohio species considered endangered, threatened, or

potentially threatened, 116 are documented as occurring in the Lake Erie

coastal zone. Specids known to occur in the coastal zone are indicated in

the species list (Appendix B). Appendix C provides a list of rare plant

species occurring in the coastal zone by county. This data should be

especially beneficial for environmental impact assessment purposes and may

be considered accurate as of January 1, 1981.

It is anticipated that the state's official lists of endangered and

threatened plants will be amended in November, 1981. In addition, field

surveys will be conducted throughout the state in 1981, and it is likely

that additional rare plant occurrences, and perhaps additional species,

will be found in the Lake Erie coastal zone in 1981.

Of the 116 species currently documented as occurring in the coastal

zone, 19 are classified as endangered, 53 as threatened, and 43 as potenti-

ally threatened. One plant is currently classified as presumed extirpated

on the state's official list. Its status will be changed when the rules

implementing O.R.C. Chapter 1518 are revised. Table I provides a breakdown

of the species totals by county.

Table I: Rare Plants of the Coastal Zone: Species Totals by County
Species Status in Ohio
(Previously)
Total # Potentially Presumed
County Species Endangered Threatened Threatened Extirpated

Ashtabula 26 2 11 12 0
Cuyahoga 14 1 4 9 0
Erie 48 1 25 22 0
Lake 15 1 4 10 0
Lorain 3 0 0 3 0
Lucas 25 3 8 13 1
Ottawa 84 14 38 31 1
Sandusky 2 1 0 1 0
Wood 0 0 0 0 0

Coastal
Zone Total 116 19 53 43 1

In the 1979 CEIP project, five specific sites were surveyed at the request

of the Coastal Energy Impact Program Administrator. Three of these areas

were either adequately surveyed in 1979 or held an extremely low possibility

of additional rare plant occurrences being found. The other two areas were

revisited in 1980, but no significant new data was obtained.

It is important to note that the data presented in this report represents

the total data base on the coastal zone and not just data collected as a

result of the 1980 CEIP project.

FEDERALLY LISTED SPECIES

The U.S. Fish and Wildlife Service currently lists (Federal Register,

Vol. 45, No. 242, December 15, 1980) one Ohio plant as threatened and fifteen

as under consideration for listing as endangered or threatened. According

to the Fish and Wildlife Service, consideration should be given to all six-

teen of these plants in environmental planning. The list is attached as

Appendix D.

Of these sixteen plants, eleven are currently known to occur in Ohio, and

five are presumed extirpated from the state. Four, and perhaps five, of these

plants historically occurred in the coastal zone, and two are currently known

to occur there. One additional plant, the Beach Sumac (Rhus trilobata var.

arenaria), may have been rediscovered as the result of the 1980 CEIP project,

but the specimens were found too late in the year for accurate identification.

These plants will be checked in May, 1981, and proper identification will be

made. The sumac is currently listed as presumed extirpated from Ohio.

The Lakeside Daisy (Hymenoxys acaulis var. glabra) and the Prairie

White-fringed Orchid (Platanthera leucophaea) are currently known to exist

in the coastal zone. The orchid was presumed extirpated from Ohio until

1980, when it was rediscovered in the western basin of the coastal zone.

Hill's Pondweed (Potamogeton hillii) historically occurred in the coastal

zone in Ashtabula County, but the site has been thoroughly altered and it is

extremely unlikely that this plant will be rediscovered. The Heart-leaved

Plantain (Plantago cordata) once occurred in Erie, Lorain, and Lucas counties,

but existing locational information is not adequate to determine whether it

was actually in the coastal zone.

MANAGEMENT CONS IDERAT IONS

All of the endangered, threatened and potentially threatened plant

species occurring in the coastal zone are significant elements of Ohio's

natural heritage and should be protected whenever possible. If priorities

must be set, the greatest protection efforts should be extended to those

species whose population levels are critically low or in greatest danger

of destruction.

It is impossible in this stage of the art to list sophisticated manage-

ment techniques appropriate for each endangered or threatened species found

in the coastal zone. Such will require detailed observations and experimental

studies of each species' needs and limitations for soil types, water quality

and quantities, microclimate conditions and extremes, effects of associated

species, pollinators, critical population sizes for outcrossing, and other

important factors. Nevertheless, much, if not most, protection may be

afforded by simply using common sense.

One generality which usually applies is that a species will be protected

best by maintaining those habitat conditions in which it currently thrives,

or at least survives. This, of course, assumes that the species is also

protected from destructive humans. The habitat conditions in which a species

thrives, however, are not always apparent. Hence, we have no good alternative

but to attempt to maintain those conditions in which we think a population

is thriving, until we know otherwise.

It will be many years before significant action is taken on most of the

identified endangered and threatened plant locations, but is is likely that

action will be taken on the several species in greatest need of protection.

In general, the Division of Natural Areas and Preserves will concentrate its

efforts on the highest quality natural areas in the coastal zone which

require additional protection.

PROTECTION STRATEGY

Although research efforts concentrated on rare plant species in 1980

as it did in 1979, significant plant communities and natural areas also

were studied as part of this project. One previously unknown area of state-

wide significance was discovered as a result of this study, and information

was collected which has aided the Division of Natural Areas and Preserves

in the continuing evaluation of coastal zone plant communities and natural

areas. There has been no significant change in the level of protection

afforded these areas since the 1979 report was developed. Significant areas

are protected by several divisions of the Ohio Department of Natural Resources,

by the U.S. Fish and Wildlife Service, by local government, and by private

organizations such as the hunting clubs of the Sandusky Bay area. There

remain several areas in the coastal zone which require additional or increased

protection. The Division will continue to concentrate preservation efforts

on the most significant areas in greatest need of additional protection.

The Division of Natural Areas and Preserves has provided some protection

to endangered and threatened plant species in the coastal zone and statewide

by implementing Ohio Revised Code Chapter 1518 (endangered plant law) on

~July 1, 1980, and by making information available through the heritage

program data services system. This system provides information to developers,

consultants,, local governments and others at a level of detail which previously

was unavailable. In addition, the DNAP has contacted some of the owners and/or

managers of significant areas to inform them of the value of the natural

features on their property. Where possible, the DNAP will assist these land-

owners in the development of management plans designed to protect the natural

features on their property. The DNAP will work with the owners of the most

significant areas in an attempt to provide the greatest level of protection

possible to these areas.

The attitude and interests of the landowner are of paramount importance

in determining the actions taken by the Division. The DNAP believes it is

essential that the landowner be the first to hear of the Division's concern

for the natural features on their property. Protection efforts generally

are more successful if the landowner is provided an accurate description of

the Division's interest from the Division rather than from a third party.

Natural area protection can take several forms, all of which have differ-

ent costs and benefits for both the landowner and the Division. Usually

some protection is provided by merely notifying the landowner of the natural

value of their property and encouraging its preservation. This, at least,

prevents the unintentional destruction of valuable natural lands. The Division

can work with landowners to develop management plans which are both compatible

with natural feature preservation and limited use by the landowner. Conser-

vation easements can provide some protection while allowing the landowner to

maintain certain rights and privileges of ownership. There are also substan-

tial tax advantages to the landowner who agrees to protect the natural value

of the property on a long term basis. Dedication as a state nature preserve

under the Ohio Natural Areas Law provides the best possible protection for

a natural area in Ohio.

The preservation technique selected by the landowner should be determined

on a case-by-case basis after exploring all alternatives. Additional infor-

mation and technical assistance can be obtained from the Division.

I1I

SUMMARY AND CONCLUSIONS

The Ohio Department of Natural Resources, Division of Natural Areas and

Preserves, by means of a planning grant from the Ohio Department of Energy,

Coastal Energy Impact Program, conducted an early summer survey of endangered,

threatened, and potentially threatened plant species; significant plant commun-

ities; and natural areas in the Lake Erie coastal zone in 1980. Data were

obtained primarily from staff field surveys.

Numerous additional records of rare plant species were identified as a

result of these field surveys. Some additional information was collected on

the relative quality of plant communities and natural areas in the coastal

zone. Surveys were conducted on several areas previously identified as requir-

ing greater protection than is currently afforded. Specific sites of concern

to the Coastal Energy Impact Program were surveyed and evaluated for early

summer flora.

Information collected via the Coastal Energy Impact Program planning

grant assisted the Division in the development of the endangerment status

for plants under Ohio Revised Code Chapter 1518 (Endangered Plant Law).

Reasonably accurate data is now available on the rare plants of the Lake

Erie coastal zone. This information should be of great value in environ-

mental impact assessment and development project planning.

Data collected during 1980 and earlier will be made available to con-

cerned entities through the Natural Heritage Program data services system.

The Ohio Department of Natural Resources, Division of Natural Areas and

Preserves, commends the Ohio Department of Energy, Coastal Energy Impact

Program, for its concern -for endangered plant species, plant communities and

natural areas in the coastal zone.

APPENDIX A

PROJECT AUTHORIZATION LETTER

James A. Rhodes Robert S. Rvan
GOVERNOR DIRECTOR
30 East Broad Street 34th Floor
Columbus, Ohio 43215
(614) 466-1805

June 12, 1980

TO: ALL ODOE/CEIP Contractors

FR: Michael Adams, Jacqueline Bird

RE: Authorization to Commence Work Under CEIP Grants

This memorandum is to serve as written permission for your agency
to begin work outlined in your Ohio Department of Energy (ODOE)
contract under the Coastal Energy Impact Program. "June 12, 1980"
will be the official commencement date entered on your contract.
To allow adequate time for project completion, the "February 28,
1981" deadline date will be extended by formal contract modification
in the near future.

In regard to invoices submitted by your agency to ODOE:

1) All invoices must be submitted using the ODOE "Request
for Payment" and "Financial Statement" forms (copies
enclosed). The Request for Payment form must bear an
original signature by an appropriate authority;

2) All invoices must have documentation of expenditures
attached, i.e., copies of time sheets, travel receipts,
subcontractor invoices, etc. No reimbursement can be
made without full documentation;

3) The first invoice should be accompanied by a statement
certifying that amountsreported as staff salaries repre-
sent the actual amounts paid to staff. ODOE should be
notified of any subsequent changes.

Be aware that all CEIP grants will be audited upon project completion.
In anticipation of this, please assure that complete and accurate
records are maintained including those regarding your required 20%
match, if applicable.

The Contractor must comply with all guidelines as published in
Office of Management and Budget Circular A-102, "Uniform Adminis-
trative Requirements for Grants-in-Aid to State and Local Governments";

cont'd...

Authorization to Commence Work

General Services Administration Federal Management Circular FMC 74-4,
"Cost Principles Applicable to Grants and Contracts with State and
Local Governments"; General Services Administration Federal Manage-
mnent Circular FMC 73-2, "Audit of Federal Operations and Programs by
Executive Branch Agencies"; and any other applicable guidelines or
regulations.

ODOE recognizes the inconveniences caused by the excessive length of
the federal approval process and is attempting to alleviate them.
Federal regulations state that OCZM review shall take 45 days. ODOE
submitted its projects application to the U.S. Office of Coastal Zone
Management/CEIP on December 15, 1979 and received approval on May 19,
1980. State encumbrance procedures were then initiated and are now
complete.

We look forward to working with you on these projects. If there are
any questions on this or any other matter pertaining to CEIP, please
contact us at 614/466-1088.

Attachments

APPENDIX B

RARE SPECIES OF NATIVE OHIO WILD PLANTS

Ohio Department of Natural Resources
DIVISION OF NATURAL AREAS & PRESERVES
Fountain Square - Columbus, Ohio 43224 - (614) 466-7803
March, 1980

RARE SPECIES OF NATIVE OHIO WILD PLANTS

The attached list represents a tabulation of Ohio endancjered, threat-
ened, potentially threatened, and presumed extirpated vascular plant
species as determined by the Division of Natural Areas and Preserves'
Natural Heritage Program on the indicated date. The list was compiled
pursuant to Ohio Revised Code Chapter 1518 which directs the Chief of the
Division of Natural Areas and Preserves to adopt criteria and lists of
plants endangered or threatened in the state. The Division gratefully
acknowledges the cooperation and assistance of the Ohio Biological Survey
Endangered Species and Population Committee in the development of this
list.

The potentially threatened and presumed extirpated species are listed
for scientific and public education purposes. Rare hybrids are included
on this list, but are not considered special plant species in the Natural
Heritage Program data base.

The species are listed by family according to classic phylogenetic
order. Nomenclature is generally that of M.L. Fernald, 1950, Gray's
Manual of Botany, eighth edition. Where more recent names are employed,
applicable synonyms in Fernald are indicated in parentheses. A column
marked "Code No." indicates data processing numbers utilized by the
Natural Heritage Program. The columns marked "US" and "OH" indicate status
levels of the species as assigned by the U.S. Fish and Wildlife Service and
by the Ohio D.N.R., Division of Natural Areas and Preserves. Status codes
are as follows:

X - Presumed Extirpated PT - Proposed Threatened (Fed. status)
E - Endangered P - Potentially threatened
PE - Proposed Endangered (Fed. status) H - Hybrid
T - Threatened

This list will be updated annually as new data become available.
Revised lists should be available in April of each year. Rules governing
endangered and threatened plants are available from the Division of Natural
Areas and Preserves.

Questions should be directed to the Natural Heritage Program in the
Division of Natural Areas and Preserves.

JAMES A. RHODES, Governor - ROBERT W. TEATER, Director - RICHARD E. MOSELEY, Chief

DESIGNATION CRITERIA

Endangered Species: A native Ohio plant species may be designated endangered
if, based on its known status in Ohio, one or more of the following criteria
applies:

(1) The species is a federal endangered species extant in Ohio.

(2) The natural populations of the species in Ohio are limited to three or
fewer sites, each of which is one square mile or less in area and is
separated from each of the other sites by at least two linear miles.

(3) The distribution of the natural populations of the species in Ohio is
limited to a geographic area delineated by one or two United States
Geological Survey 7.5 minute quadrangle maps.

(4) The total number of plants in all the natural populations of the species
in Ohio is limited to one hundred or fewer individual, physically
unconnected plants.

Threatened Species: A native Ohio plant species may be designated threatened
if, based on its known status in Ohio, one or more of the following criteria
applies:

(1) The species is a federal threatened species extant in Ohio but not on the
state endangered species list.

(2) The natural populations of the species in Ohio are limited to no less than
four nor more than ten sites, each of which is one square mile or less in.
area and is separated from each of the other sites by at least two linear
miles.

(3) The distribution of the natural populations of the species in Ohio is
limited to a geographic area delineated by no less than three nor more
than five United States Geological Survey 7.5 minute quadrangle maps.

Presumed Extirpated Species: A native Ohio plant species may be designated
presumed extirpated when no natural populations of the species are known to
occur in the state.

Potentially Threatened Species: A native Ohio plant species may be designated
potentially threatened if it qualifies for one or more of the following criteria.

(1) The species is extant in Ohio and does not qualify as a state endangered
or threatened species, but it is either
a. a proposed federal endangered or threatened species or a species
listed in the Federal Register as under review for such proposal,
and/or

b. a species protected under the Convention on International Trade in
Endangered Species of Wild Fauna and Flora.

(2) The natural populations of the species are imperiled to the extent that
the species could conceivably become a threatened species in Ohio within
the foreseeable future.

(3) The natural populations of the species, even though they are not threatened
in Ohio at the time of designation, are declining in abundance or vitality
at a significant rate throughout all or large portions of the state.
16

March, 1980

DIVISION OF NATURAL AREAS AND PRESERVES
NATURAL HERITAGE PROGRAM
Special Plants

Pteridophytes COASTAL
ZONE
Code No. COUNTIES OH
LYCOPODIACEAE. Clubmoss Family
Lycopodium inundatum, Bog Clubmoss .......................EA F46 P

SELAGINELLACEAE. Spikemoss Family
Selaginella rupestris, Rock Spikemoss ....................EY F47 T

ISOETACEAE. Quillwort Family
Isoetes engelmannii (I. engelmanni), Appalachian
Quillwort ........................................... EB 359 E
Isoetes muricata, Spiny-spored Quillwort .................EB F48 X

EQUISETACEAE. Horsetail Family
Equisetum sylvaticum, Woodland Horsetail .................EZ A07 T
Equisetum variegatum (E. variegatum var. variegatum),
Variegated Scouring-rush ............................ EZ F50 0 P
Equisetum X nelsonii (E. variegatum var. nelsoni),
Nelson's Scouring-rush .........EZ F51 H
Equisetum X trachyodon (E. variegatum var. jesupi),
Jesup's Scouring-rush .......................... EZ F52 H

OPHIOGLOSSACEAE. Adder's-tongue Family
Botrychium biternatum (B. dissectum var. tenuifolium;
see Gleason and Cronquist), Sparse-lobe Grape-fern..EC F53 E
Botrychium lanceolatum, Triangle Grape-fern ..............EC 082 E
Botrychium multifidum, Leathery Grape-fern ... ............ EC F54 T
Botrychium simplex (including varieties & formae),
Least Grape-fern ....................................EC F55 E
Botrychium ternatum (not in Fernald 1950; see Wherry
1961), American Ternate Grape-fern ... ...............EC F56 E
Ophioglossum engelmannii (O. engelmanni), Limestone
Adder's-tongue ..................................... EC F57 E

SCHIZAEACEAE. Curly-grass Family
Lygodium palmatum, Climbing Fern .........................ED 429 P

ADIANTACEAE (POLYPODIACEAE). Maidenhair Family

Vittaria lineata (not in Fernald 1950; see Wagner &
Sharp, 1963), Appalachian Gametophyte ...............EF B08 T

HYMENOPHYLLACEAE. Filmy Fern Family
Trichomanes boschianum, Appalachian Filmy Fern ...........EE 726 E

-2-

Code No. OH
POLYPODIACEAE. Polypody Family
Polypodium polypodioides, Little Gray Polypody.......EF F69 E

ASPLENIACEAE ( POLYPODIACEAE). Spleenwort Family
Asplenium bradleyi, Bradley's Spleenwort..........EF 0,57 E
Asplenium montanum, Mountain Spleenwort ....... . ..EF F58 P
Asplenium resiliens, Black-stem Spleenwort.........EF 063 X
Asplenium ruta-muraria var. cryptolepis (A. cryptolepis
var. cryptolepis & var. ohionis; including A. r.
var. subtenuifolium), Wall-rue ....... . ....EF 065 T
Asplenium X clermontae (not In Fernald 1950; see
Wherry 1961), Miss Clermont's Spleenwort .......EF F59 H
Asplenium X ebenoides (X Asplenosorus ebenoides),
Scott's Spleenwort....................EFF60 H
Asplenium X inexpectatum (X Asplenosorus inexpectatus),
Unexpected Spleenwort . .................EFF61 H
Asplenium X kentuckiense (not in Fernald 1950; see
Wherry 1961 under A. X Kentuckyense), Kentucky
S pleenwort .. .........................EF F62 H
Asplenium X trudellii (X A. trudelli), Trudell's
Spleenwort .......................EF 066 H
Cystopteris X tennesseensis (not in Fernald 1950; see
Wherry 1961), Tennessee Bladder Fern ..........EF 209 H
Dryopteris clintoniana (D. cristata var. clintoniana),
Clinton's Wood Fern ....................EF F64 E
Dryopteris X boottil (X D. boottii), Boott's Wood Fern...EF F65 H
Dryopteris X pittsfordensis (not in Fernald 1950; see
Wherry 1961), Pittsford Wood Fern....... . ...EF BIO H
Dryopteris X slossonae (not in Fernald 1950; see
Wherry 1961), Miss Slosson' s Wood Fern ... ......EF F66 H
Dryopteris clintoniana (D. cristata var. clintoniana)
X Dryopteris glia...q... ............*.. *EF F67 H
Dryopteris goldiana X Dryopteris intermedia .............. EF F68 H

Gymnocarpium dryopteris (Dryopteris disjuncta), Oak
Fern ..........................EF 296 T

Phegopteris connectilis (Thelypteris phegopteris;
Dryopteris phegopteris), Long Beech-fern .......EF 720 P
Woodsia ilvensis, Rusty Woodsia ...... . .......EF759 X

BLECHNACEAE (POLYPODIACEAE). Blechnurn Family
Anchistea virginica (Woodwardia virginica), Virginia
Chain-fern.. . .. .. ................... .EF 761 P
Lorinseria areolata (Woodwardia areolata), Netted
Chain-fern . ........ ..............EF F70 E

Gymnosperms

PINACEAE. Pine Family
Larix laricina, Tamarack ....... . .........FA 377 P
Pirnus echinata, Yellow Pine....... . ........FA 775 P

CUPRESSACEAE (PINACEAE). Cypress Family
Juniperus comumunis (including varieties) Ground Juniper. FC 371 T
Thuja occidentalis, Arbor Vitae..............FC 776 0 P
18

Angiosperms - Monocotyledons

Code No. OH

SPARGANIACEAE. Bur-reed Family
Sparganium androcladum, Keeled Bur-reed ...........GB A70 Lt, T
Sparganium chlorocarpum, Small Bur-reed ....... . ..GB 686 E

POTAMOGETONACEAE (ZOSTERACEAE). Pondweed Family
Potamogeton amplifolius, Large-leaved Pondweed.......GC 543 p
Potamogeton epihydrus, Nuttall's Pondweed .........fc F25 A P
Potamogeton filiformis, Filiform Pondweed .........GC H51 0 E
Potamogeton friesii, Fries' Pondweed....... . ....GC H52 X
Potamogeton gramineus, Grass-like Pondweed.........GC H53 E
Potamogeton hillii, Hill's Pondweed.............GC H54 X
Potamogeton natans, Floating Pondweed ...........GC 547 O, Lo 0Lu T
Potamogeton perfoliatus, Red-head Pondweed.........GC H55 X
Potamogeton praelongus, White-stem Pondweed ........GC H56 E
Potamogeton pulcher, Spotted Pondweed ....... . ...GC B16 E
Potamogeton richardsonii, Richardsc'n's'Pondweed ......GC H58 AQ0 T
Potamogeton robbinsii, Robbins' Pondweed....... . ..GC H59 E
Potamogeton spirillus, Spiral Pondweed .. ...... . ..GC B17 T
Potamogeton strictifolius, Straight-leaved Pondweed ... ... .GC H60 X
Potamogeton tennesseensis, Tennessee Pondweed .......GC 549 E
Potamogeton vaseyi, Vasey's Pondweed......... . ..GC H-61 X
Potamogeton zosteriformis, Flat-stem Pondweed ..........GC B18 0 T

NAJADACEAE. Naiad Family
Najas flexilis, Slender Naiad.................GZ E95 AOE P
Najas gracillima, Thread-like Naiad ...... . .....GZE96 X

JUNCAGINACEAE. Arrow-grass Family
Scheuchzeria palustris, Scheuchzeria ......... . ...GE 634 E
Triglochin maritimum (T. maritima), Seaside Arrow-grass..GE H62 T.
Triglochin palustre (T. palustris), Marsh Arrow-grass .... GE H63 T

ALISMATACEAE. Water-Pl antain Family
Echinodorus rostratus, Bur-head.. ............ .GF H64 T
Lophotocarpus calycinus, Southern Wapato ............GF A72 EO P
Sagittaria australis, Long-beaked Arrowhead . ............GF H65 'IT
Sagittaria cuneata, Wapato ....... ................GF H66 EOL'- T
Sagittaria graminea, Gras�-Ieaf Arrowhead ..........GF 618 0T
Sagittaria latifolia var. pubescens, Hairy Arrowhead.....GF H67 E

HYDROCHARITACEAE. Frog's-bit Family

Vallisneria americana, Eel-grass ..............GG H68 AILo..,C.*6,o P

-4-

Code No. OH

GRAMINEAE (P0ACEAE). Grass Family
Agropyron trachycaulum, Bearded Wheat Grass .........GH 015 0P
Agrostis elliottiana, Elliott's Bent-grass .........GH 017 E
Animophila breviligulata, American Beach Grass .......GH H69/ All_0_C.,EQL~AT
Aristida purpurascens, Purple Triple-awined Grass . . ....GH H70 T
Calamagrostis inexpansa, Northern Reed-grass . . ......GH H71 E
Calamagrostis insperata (not in Fernald; see Braun),
Bartley's Reed Bent Grass....... . .......GH H72 E
Cinna latifolia, Northern Wood-reed ....... . ....GH H74 E
Danthonia compressa, Flattened Wild Oat Grass . . ......GH H75 T
Deschampsia caespitosa, Tufted Flairgrass .. ...... . ..GH 214 E
Deschampsia flexuosa, Crinkled Hairgrass ........ ..GH H76 T
Digitaria filiformis, Slender Finger-grass . . .......GH B26 E
Erianthus alopecuroides, Silver Plume Grass .........GH H77 E
Glyceria acutiflora, Sharp-glumed Manna-grass . . ......GH B28 E
Glyceria borealis, Northern Manna-grass........ . ..GH 878 X
Glyceria grandis, Tall Manna-grass ........ . ....GH B30 T
Glyceria melicaria, Long Manna-grass ........ . ...GH H79 P
Glyceria pallida, Pale Manna-grass ........ . ....GH 292 p
Gymnopogon ambiguus, Beardgrass ....... . ......GH H80 E
Koeleria macrantha (K. cristata), Junegrass.........GH HSI T
Leptoloma cognatum, Fall Witch-gSrass ............GH H82 C- T
Melica nitens, Three-flowered Melic ........ . ....GH 446 E
Muhlenbergia capillaris, Hairgrass....... . .....GH H83 X
Muhlenbergia curtisetosa, Short-awned ~!irnblewjll . . ....GH H84 X
Muhlenbergia cuspidata, Plains Muhlenbergia.........GH H85 L',.. E
Panicum bicknellii, Bicknell's Panic-arass . . ..........GH H86 X
Panicum boreale, Northern Panic-grass........ . ...GH H87 E
Panicum calliphyllum, Tall Green Panic-grass . . ......GH H88 X
Panicum columbianum, American Panic-grass . ....... ..GH B39 T
Panicum depauperatum, Starved Panic-grass . ...........GH H89 P
Panicum laxiflorum, Pale Green Panic-grass .. .......GH H90 T
Panicum leibergii, Leiberg's Panic-grass ........ ..GH H91 X
Panicumn longifolium, Long-leaved Panic-grass . . ......GH H92 X
Panicum oligosanthes, Prairie Panic-grass. ....... ..GH 893 P
Panicum philadelphicum, Philadelphia Panic-grass .......GH 488 P
Panicum verrucosum, Warty Panic-grass..... ..........GH H94 E
Panicum villosissimum, Villous Panic-grass .........GH H95 E
Panicum yadkinense, Spotted Panic-prass . ....... . ..GH B41 X
Paspalum setaceum, Slender Paspalum........ . ....GH H96 X
Poa languida, Weak Spear-grass . ....... ..........GH 529 T
Poa paludigena, Marsh Spear-grass . .............GH H97 X
Poa wolfii, Wolf's Bluegrass .................GH H98 X
Schizachne purpurascens, False Melic ........ . ...GH 846 E
Sphenopholis obtusata, Prairie Wedgegrass ...........GH B48 T
Sporobolus cryptandrus, Sand Dropseed . . . G................. GH H99P
Sporobolus heterolepis, Prairie Dropseed ........ ..GH 693 E
Stipa avenacea, Blackseed Needle Grass........ . ..GH B49 E
Stipa spartea, Porcupine Grass ....................GH G67 E
Triplasis purpurea, Purple Sand-grass ..... .............GH JOO ~L.,oE P
0I-A

-5-

Code No. OH

GRAMINEAE (POACEAE). Grass Family (Cont'd.)
Trisetum pensylvanicum, Swamp-oats ...................... GH J01 T
Uniola latifolia, Wild Oats ............................. GH J02 P
Zizania aquatica, Wild Rice ............................. GH B03 0 T

CYPERACEAE. Sedge Family
Carex aggregata, Crowded Sedge .......................... GJ B50 P
Carex alata, Broad-winged Sedge ......................... GJ J03 E
Carex albolutescens, Pale Straw Sedge ................... GJ J04 E
Carex amphibola var. rigida, Stiff Wood Sedge ........... GJ J05 E
Carex annectens var. xanthocarpa, Yellow-fruited Sedge.. GJ R57 T
Carex aquatilis, Leafy Tussock Sedge .................... GJ J06 ALto E
Carex arctata, Drooping Wood Sedge ...................... GJ J07 E
Carex argyrantha, Silvery Sedge ......................... GJ J08 E
Carex atherodes, Wheat Sedge ............................ GJ A14 � T
Carex aurea, Golden-fruited Sedge ....................... GJ J09 A,0 T
Carex baileyi, Bailey's Sedge ........................... GJ 110 X
Carex bebbii, Bebb's Sedge ............ .................. GJ J10 E
Carex brevior, Tufted Fescue Sedge ...................... GJ Jll 0 P
Carex brunnescens, Brownish Sedge ....................... GJ J12 X
Carex buxbaumii, Brown Bog Sedge .........................GJ 113 P
Carex cephalantha, Little Prickly Sedge ................. GJ J13 E
Carex cephaloidea, Thin-leaf Sedge ...................... GJ J14 E
Carex conoidea, Field Sedge ............................. GJ 118 T
Carex crawei, Crawe's Sedge ............................. GJ E33 0 T
Carex crus-corvi, Raven-foot Sedge ...................... GJ J15 X
Carex cryptolepis (C. flava var. fertilis), Little
Yellow Sedge ....................................... GJ J16 P
Carex debilis var. debilis, Weak Sedge .................. GJ J17 E
Carex debilis var. rudgei, Rudge's Sedge ................ GJ J17 P
Carex decomposita, Cypress-knee Sedge ................... GJ 120 X
Carex deweyana, Dewey's Sedge ........................... GJ J19 X
Carex diandra, Lesser Panicled Sedge .................... GJ J20 E
Carex eburnea, Bristle-leaf Sedge ....................... GJ 122 C,0 P
Carex emmonsii, Emmons' Sedge ........................... GJ J21 E
Carex festucaceao- Fescue Sedge ......................... GJ J22 P
Carex flava (C. flava var. flava, but not C. flava var.
fertilis. See C. cryptolepis.), Yellouw Sedge ...... GJ J23 E
Carex foenea, Hay Sedge ............. .................... GJ J24 X
Carex folliculata, Long Sedge ........................... GJ J25 T
Carex formosa, Handsome Sedge ........................... GJ J26 E
Carex glaucodea (C. flaccosperma var. glaucodea), Blue-
green Sedge ........................................ GJ J27 T
Carex gravida, Heavy Sedge .............................. GJ E34 T
Carex haydenii, Hayden's Sedge .......................... GJ J28 E
Carex howei, Howe's Sedge ............................... GJ B57 E
Carex incomperta, Prickly Bog Sedge ..................... GJ J29 P
Carex interior, Inland Sedge ............ ................ GJ J30 T
Carex lasiocarpa, Slender Sedge ......................... GJ A15 � T
Carex limosa, Mud Sedge ................................. GJ J31 X
Carex louisianica, Louisiana Sedge ...................... GJ J32 E
Carex lupuliformis, False Hop Sedge ..................... GJ 126 0 P

Code No. OH

CYPERACEAE. Sedge Family (Cont'd.)
Carex meadii, Mlead's Sedge .................GJ B61 T
Carex merritt-fernaldii, Fernald's Sedge ..........GJ J33 X
Carex mesochorea, Midland Sedge . ..............GJ B64 X
Carex muhlenbergii, Muhlenberg's Sedge ...........GJ J34 0P
Carex nigromarginata, Black-margined Sedge .........GJ B66 E
Carex oligosperma, Few-seeded Sedge . ............GJ 335 E
Carex ormnostachya, Stiff Broad-leaved Sedge .........GJ L56 E
Carex pallescens, Pale Sedge . ....:...........GJ3J36T
Carex pedunculata, Long-stalked Sedge ............G3 B67 T
Carex prairea, Prairie Sedge ................GJ A16 P
Carex projecta, Necklace Sedge ...............GJ 130 T
Carex pseudo-cyperus, Smaller Bearded Sedge . ........GJ J37 X
Carex radiata, Radiate Sedge ...................GJJ38 T
Carex retroflexa, Reflexed Sedge ..............GJ J39 T
Carex retrorsa, Reflexed Bladder Sedge ...........GJ J40 X
Carex richardsonii, Richardson's Sedge . ........ ...GJ J41 X
Carex rostrata, Beaked Sedge ................GJ E39 T
Carex sartwellii, Sartwell's Sedge .............GJJ42 T
Carex scabrata, Rough Sedge . ................GJ J43 T
Carex seorsa, Weak Bog Sedge ................GJ J44 T
Carex sprengelii, Sprengel's Sedge .............GJJ45 Q E
Carex sterilis, Fen Sedge . ..................GJ 46 E
Carex straminea, Straw Sedge ................GJ 47 E
Carex striatula, Lined Sedge . ........ ..........GJ G83 X
Carex suberecta, Prairie Straw Sedge ............GJ B70 E
Carex Xsullivantii, Sullivant's Sedge ......... . ..GJ R58 H
Carex tenera, Slender Straw Sedge . .............GJ J49 P
Carex tetanica, Slender Millet Sedge ............GJ 135 T
Carex trichocarpa, Hairy-fruited Sedge ...........GJ B72 E P
Carex trisperma, Three-seeded Sedge........ .....GJ 137 T
Carex viridula, Little Green Sedge ...............GJ J50 EjO T
Carex willdenowii, Willdenow's Sedge ............GJ J51 T
Cladium mariscoides, Twig-rush . ...............GJ3E41 0 T
Cyperus acuminatus, Pale Umbrella-sedge...........GJ J52 E
Cyperus diandrus, Low Umbrella-sedge ...............GJ J53 E,1t.. T
Cyperus dipsaciformis, Teasel-sedge.. ..... .........GJ J54 E
Cyperus engelmannii (C. engelmanni), Engelmann' s
Umbrel la-sedge. . ... . ... . .. . .. ....... ........... GJ E42 EIc', Lu_. P
Cyperus houghtonii, Houghton's Umbrella-sedge.. ..... .GJ J55 X
Cyperus lancastriensis, Many-flowered LUmbrella-sedge ..... GJ J56 T
Cyperus refractus, Reflexed Umnbrella-sedge . . ........GJ B75 T
Cyperus schweinitzii, Schweinitz's Umbrella-sedge ........ GJ J57 NC 1I" P
Eleocharis compressa, Flat-stem Spikerush . . ............GJ B77 EJO)L~c P
Eleocharis elliptica, Yellow-seeded Spikeru-sh.. . .......GJ J58 E: T
Eleocharis intermedia, Matted Spikerush .................. GJ A17 Of-,L- T
Eleocharis olivacea, Olivaceous Spikerus h ...........GJ 359 0T
Eleocharis pauciflora, Few-flowered Spikerush ............ GJ J60 E
Eleocharis quadrangulata, Four-angled Spikerush ........ .G J61 T
Eleocharis rostellata, Beaked Spikerush .................. GJ 247 P
Eleocharis wolfii, Wolf's Spikerush.. .... ............. GJ 62, X
Eriophorum gracile, Slender Cottongrass...... . ... GJ J63 X

-7-

Code No. OH

CYPERACEAE. Sedge Family (Cont'd.)
Eriophorum virginicum, Tawny Cottongrass ................ GJ E48 T
Eriophorum viridi-carinatum, Green Cottongrass . ......... GJ J64 T
Hemicarpha micrantha, Dwarf Bulrush ..................... GJ 317 E
Rhynchospora alba, White Beak-rush ...................... GJ 600 T
Rhynchospora globularis, Grass-like Beak-rush ........... GJ B78 E
Scirpus expansus, Woodland Bulrush ...................... GJ E56 E

Scirpus purshianus, Pursh's Bulrush ..................... GJ B79 0 T
Scirpus smithii, Smith's Bulrush ........................ GJ J65 0 E
Scirpus subterminalis, Swaying Rush ..................... GJ 646 E
Scirpus torreyi, Torrey's Bulrush ....................... GJ J66 X
Scleria pauciflora, Few-flowered Nut-rush ............... GJ J67 T
Scleria triglomerata, Tall Nut-rush ..................... GJ B82 T
Scleria verticillata, Low Nut-rush ...................... GJ E58 E T

ARACEAE. Arum Family

Arisaema stewardsonii, Swamp Jack-in-the-pulpit ......... GL J68 P
Arisaema triphyllum, Small Jack-in-the-pulpit ........... GL J69 P
Calla palustris, Wild Calla ............................. GL J70 P

LEMNACEAE. Duckweed Family

Lemna valdiviana, Pale Duckweed ......................... GM B83 X
Wolffia papulifera, Pointed Water-meal .................. GM 758 T
Wolffiella floridana, Wolffiella ........................ GM J71 X

XYRIDACEAE. Yellow-eyed-grass Family
Xyris difformis (X. caroliniana), Carolina Yellow-eyed-
grass .............................................. GN J72 X
Xyris torta, Twisted Yellqw-eyed-grass .................. GN 766 E

ERIOCAULACEAE. Pipewort Family
Eriocaulon septangulare, White-buttons .................. GP 259 X

PONTEDERIACEAE. Pickerelweed Family
Heteranthera reniformis, Mud-plantain ................... GR A20 X

JUNCACEAE. Rush Family
Juncus alpinus, Alpine Rush ............................. GS J74 AE,O T
Juncus balticus, Baltic Rush ............................ GS 367 ,0O T
Juncus greenei, Greene's Rush ........................... GS J75 T
Juncus interior, Inland Rush ............................ GS J76 0,S E
Juncus platyphyllus, Flat-leaved Rush ................... GS B88 T
Juncus scirpoides, False Bulrush ........................ GS B89 E T
Juncus secundus, One-sided Rush ......................... GS J77 E T
Luzula bulbosa, Southern Woodrush ....................... GS B91 T

Code No. OH

LILIACEAE. Lily Family
Aletris farinosa, Unicorn-root ...........................GT J79 P
Clintonia borealis, Bluebead-lily ........................GT E85 E
Clintonia umbellulata, Speckled Wood-lily ................GT J80 T
Disporum maculatum, Nodding Mandarin .....................GT 223 T
Erythronium rostratum (not in Fernald 1950; see Braun
1967), Goldenstar . .........GT E86 E
Lilium philadelphicum, Wood-lily . ........................GT 398 O P
Lilium superbum, Turk's-cap Lily .......... ...............GT 769 P
Melanthium virginicum, Bunchflower . ......................GT E90 T
Nothoscordum bivalve, False Garlic . ......................GT J81 E
Smilacina trifolia, Three-leaved Solomon's-seal ..........GT J82 X
Smilax illinoensis, Illinoian Smilax .....................GT J83 H
Smilax lasioneura, Pale Carrion-flower . .................. G J84 P
Smilax pulverulenta, Downy Carrion-flower . ...............GT J85 E
Stenanthium gramineum, Feather-bells . ....................GT J86 P
Streptopus roseus, Rose Twisted-stalk . ...................GT 704 E
Tofieldia glutinosa, False Asphodel . .....................GT A24 T
Trillium cernuum, Nodding Trillium . ......................GT 732 X
Trillium nivale, Snow Trillium ......... . .................GT A74 P
Trillium recurvatum, Prairie Wake-robin . .................GT J87 E
Trillium undulatum, Painted Trillium . ....................GT J88 E
Veratrum viride, White Hellebore .........................GT J89 P
Veratrum woodii, Wood's Hellebore . .......................GT E93 T
Zygadenus glaucus (Zigadenus glaucus), Wand-lily .........GT 767 T

AMIARYLLIDACEAE. Amaryllis Family
Agave virginica, American Aloe ............... ............GV A25 P

IRIDACEAE. Iris Family
Iris brevicaulis, Leafy Blue Flag ........................GX J91 E E
Iris verna, Dwarf Iris ........... ........................GX 357 E
Sisyrinchium atlanticum, Atlantic Blue-eyed-grass ........GX J92 E
Sisyrinchium montanum, Northern Blue-eyed-grass ..........GX J93 E T
Sisyrinchium mucronatum, Narrow-leaved Blue-eyed-grass...GX J94 E10 T

-9-

Code No. OH

ORCHIDACEAE. Orchid Family

Arethusa bulbosa, Dragon's-mouth ........................ GY 051 E
Calopogon tuberosus (C. pulchellus), Grass-pink ......... GY K69 P
Coeloglossum viride (Habenaria viridis), Long-bracted
Orchid ............................................. GY A28 E
Corallorhiza maculata, Spotted Coral-root ............... GY 177 P
Corallorhiza trifida, Early Coral-root .................. GY A27 E
Corallorhiza wisteriana, Spring Coral-root .............. GY 179 E
Cypripedium acaule, Stemless Lady's-slipper ............. GY 204 P
Cypripedium calceolus var. parviflorum, Small Yellow
Lady's-slipper ..................................... GY 205 E
Cypripedium calceolus var. pubescens, Large Yellow
Lady's-slipper ..................................... GY 205 P
Cypripedium candidum, White Lady's-slipper .............. GY K52 E
Cypripedium reginae, Showy Lady's-slipper ............... GY 206 T
Goodyera tesselata, Checkered Rattlesnake-plantain ...... GY K53 X
Hexalectris spicata, Crested Coral-root ................. GY 328 E
Isotria verticillata, Whorled Pogonia ................... GY 364 P
Liparis loeselii, Fen Orchid ............................ GY 404 A,EO P
Listera cordata, Heartleaf Twayblade .................... GY 406 X
Malaxis unifolia, Green Adder's-mouth ................... GY 439 P
Platanthera blephariglottis (Habenaria blephariglottis),
White Fringed Orchid ............................... GY 298 X
Platanthera ciliaris (Habenaria ciliaris), Yellow
Fringed Orchid .................................... GY 299 T
Platanthera clavellata (Habenaria clavellata), Green
Woodland Orchid .................................... GY G95 T
Platanthera flava (Habenaria flava), Tubercled Rein-
orchid ......................................... GY 300 P
Platanthera grandiflora (Habenaria psycodes var.
grandiflora; H. fimbriata), Large Purple-fringed
Orchid ..................................... GY AC9 X
Platanthera hookeri (Habenaria hookeri), Hooker's
Orchid ..................................... GY K55 X
Platanthera hyperborea (Habenaria hyperborea), Tall
Northern Green Orchid .............................. GY K56 X
Platanthera leucophaea (Habenaria leucophaea), Prairie
Fringed Orchid .................................... GY K57 L%, O X
Platanthera orbiculata (Habenaria orbiculata), Large
Round-leaved Orchid ............................... GY 302 T
Platanthera peramoena (Habenaria peramoena), Purple
Fringeless Orchid ................................. GY 303 P
Platanthera psycodes (Habenaria psycodes var. psycodes),
Small Purple Fringed Orchid ........................ GY F05 T
Pogonia ophioglossoides, Rose Pogonia ....................GY K61 T
Spiranthes lacera var. gracilis (Spiranthes gracilis),
Southern Slender Ladies'-tresses ....................GY K63 P
Spiranthes lucida, Shining Ladies'-tresses . ..............GY A76 T
Spiranthes magnicamporum (not in Fernald 1950; see
Luer 1975), Great Plains Ladies'-tresses ............GY V12 E,O T
Spiranthes ovalis, Lesser Ladies'-tresses .................GY 691 T
Spiranthes romanzoffiana, Hooded Ladies'-tresses .........GY K65 E
Spiranthes tuberosa, Little Ladies'-tresses . .............GY A75 P

Triphora trianthophora, Three-birds-orchid ...............GY 736 T
25

-10-

Angiosperms - Dicotyledons

Code No. OH

SALICACEAE. Willow Family
Populus balsamifera, Balsam Poplar . ......................KM 777 0 E
Populus heterophylla, Swamp Cottonwood ....... ............KM 778 X
Salix bebbiana, Long-beaked Willow . . .....................KM 779 0,L- P
Salix candida, Hoary Willow . . ..............KM 780 T
Salix caroliniana, Carolina Willow .......................KM 781 E
Salix glaucophylloides var. glaucophylloides (S. glauco-
phylloides var. glaucophylla), Blue-leaved Willow...KM 782 X
Salix glaucophylloides var. albovestita, Dune Willow .....KM 782 X
Salix pedicellaris, Bog Willow . ..........................KM 783 E
Salix petiolaris (S. gracilis var. textoris), Slender
Willow ..............................................KM 784 T
Salix serissima, Autumn Willow ............. ..............KM 785 T
Salix X subsericea .......................................KM 786 H
Salix syrticola, Sand-dune Willow . .......................KM 787 X

M1YRICACEAE. Bayberry Family
Comptonia peregrina, Sweet-fern ..........................JH 788 T
Myrica pensylvanica, Bayberry .... ........................JH 789 E

BETULACEAE (CORYLACEAE). Birch Family
Betula lenta, Sweet Birch ................................JK C50 P
Betula populifolia, Gray Birch ......... . .................JK K04 P
Betula pumila, Swamp Birch ...............................JK 791 T
Corylus cornuta, Beaked Hazel .......... ..................JL 792 E

FAGACEAE. Beech Family
Castanea dentata, American Chestnut ...... ................JJ 142 P
Quercus ellipsoidalis, Jack Oak ..........................JJ 794 X
Quercus falcata, Spanish Oak ...............................JJ 795 T
Quercus marilandica, Blackjack Oak .. .....................JJ 796 T
Quercus shumardii, Shumard's Red Oak .....................JJ B93 P

ULMACEAE. Elm Family
Celtis tenuifolia, Dwarf Hackberry ....................... JD C59 � P
Ulmus americana, American Elm .................. ........... JD 797 P
Ulmus thomasii, Rock Elm .................................JD 798 E

URTICACEAE. Nettle Family
Pilea fontana, Coolwort ...................................JF 799 P
Urtica chamaedryoides, Spring Nettle .....................JF 800 E

VISCACEAE (LORANTHACEAE). Mistletoe Family
Phoradendron serotinum (P. flavescens), American
Mistletoe ..............................................MA X61 T

POLYGONACEAE. Buckwheat Family
Polygonum careyi, Carey's Smartweed ...................... JT 801 X

Code No. OH
POLYGONACEAE. Buckwheat Family (Cont'd.)
Polygonum cilinode, Mountain Bindweed ....... . ...JT 802 E
Polygonum pensylvanicum var. eglandulosum, Lake Erie
P inkweed..................... .....3T 803 E.1c T
Polygonum ramosissimum, Bushy Knotweed......... ..JT 804 0 E

CHENOPODIACEAE. Goosefoot Family
Chenopodium capitatum, Strawberry-blite ..........JQ 805 X
Chenopodium leptophyllum, Slender Goosefoot ......... Q 806 0 E

AMARANTHACEAE. Amaranth Family
Froelichia floridana, Cottonweed ......... . .....JR 807 E

PORTULACACEAE. Purslane Family
Claytonia caroliniana, Carolina Spring-beauty .......JP 808 A~T

CARYOPHYLLACEAE. Pink Family
Arenaria patula, Spreading S andwort . ...........JN 809 E
Arenaria stricta (including varieties), Rock Sandwort .... JN V96 0T
Sagina decumbens, Southern Pearlwort ................JN 810 X
Silene caroliniana var. pensylvanica, Carolina Catchfly..JN 811 T
Silene caroliniana var. wherryi, Wherry's Catchfly ....... JN 811 E
Silene nivea, Snowy Campion................ .......3N 812 X
Silene regia, Royal Catchfly ............................. JN 669 E
Silene rotundifolia, Round-leaved Catchfly ............... JN 670 T

CERATOPHYLLACEAE. Hornwort Family
Ceratophyllum echinatum, Prickly Hornwort ................ HG 813 0T

NYMPHAEACEAE. Water-lily Family
Nuphar variegatum, Bullhead Lily ................HF 814 0~" E

RANUNCULACEAE. Buttercup Family
Aconitum noveboracense, Northern flonkshocd. ... .....HH 815 E
Actaea rubra, Red Baneberry ............. .......... :::.. HH 816 E
Anemone cylindrica, Prairie Thimbleweed .................. HH 817 LUjc) T
Clematis occidentalis, Purple Virgin's-bower .. .........HH 169 X
(C. verticillaris. See Pringle 1971)
Coptis groenlandica, Goldthread .......................... HH 818 P
Delphinium exaltatum, Tall Larkspur ...................... HH 212 T
Ranunculus fascicularis, Early Buttercup .............HH 819 F-,O T
Ranunculus pusillus, Low Spearwort ....................... HH 820 E
Trollius laxus, Spreading Globe-flower ................HH 821 E

MAGNOLIACEAE. Magnolia Family
Magnolia macrophylla, Bigleaf Magnolia ............HA 822 E
Magnolia tripetala, Umbrella Magnolia .................... HA 436 T

CALYCANTHACEAE. Calycanthus, Family
Calycanthus fertilis, Sweet-shrub ........................ HC 823 E

FUMARIACEAE (PAPAVERACEAE). Fumitory Family
Adlumia fungosa, Mountain-fringe ......................... JA 006 E
Corydalis sempervirens, Rock-harlequin ................... JA 186 T

-12-
Code No. OH
CRUCIFERAE (BRASSICACEAE). Mustard Family (Cont'd.)
Arabis divaricarpa, Limestone Rock-cress ........ ....KP 824 LI... E
Arabis drummondii, Drummiond's Rock-cress ...............KP 825 0 E
Arabis hirsuta var. pycnocarpa, Western Hairy Rock-cress.KP 826 0 T
Arabis hirsuta var. adpressipilis, Southern Hairy Rock-
cress ..... *.*. ...... . . ...... .. *....... *.~ ..*.*.. .KP 826 P
Arabis lyrata, Lyre-leaf Rock-cress ...................... KP 044 La.P
Arabis patens, Spreading Rock-cress . . . .............KP 045 E
Armoracia aquatica, Lake-cress ...... ....... o .....KP 827 X
Cakile edentula var. lacustris, Inland Sea-rocket ........ KP C61 PL)C..,,P
Cardamnine rotundifolia, Mountain-watercress .............. KP 106 p
Dentaria multifida, Narrow-leaved Toothwort .............. KP 829 T
Descurainia pinnata, Tansy-mustard.............KP 830 O.SP
Draba brachycarpa, Little Whitlow-grass .................. KP 831 E
Draba cuneifolia, Wedge-leaf Whitlow-grass ............... KP 832 E
Draba reptans, Carolina Whitlow-grass .................... KP 228 T
Erysimunm arkansanuni, Western Wall-flower ................. KP 833 E
Leavenworthia uniflora, Michaux's Leavenworthia .......... KP 834 T
Rorippa sessiliflora, Sessile Yellow-dress ............... KP 835 T

SARRACENIACEAE. Pitcher-plant Family
Sarracenia purpurea, Pitcher-plant. ...... . .....KE 627 p

DROSERACEAE. Sundew Family
Drosera intermedia, Soath'ilate-leaved Sundew .............. KF 836 E
Drosera rotundifolia, Round-leaved Sundew ................ KF 231 P

SAXIFRAGACEAE. Saxifrage Family
Heuchera parviflora, Small-flowered Alumroot ............. LB 326 T
Heuchera villosa, Hairy Alumroot ......................... LB 838 T
Parnassia glauca, Grass-of-Parnassus ..................... LB C62 P
Ribes glandulosum, Skunk Currant ......................... LB 840 X
Ribes hirtellum, Smooth Gooseberry ....................... LB 841 p
Ribes lacustre, Bristly Black Currant .................... LB 842 X
Ribes missouriense, Missouri Gooseberry .................. LB 843 X
Ribes rotundifolium, Appalachian Gooseberry .............. LB 844 E
Sullivantia sullivantii, Sullivantia ..................... LB 845 p

ROSACEAE. Rose Family
Agrimonia striata, Hairy Agrimony ........................ LC 846 T
Amnelanchier laevis, Allegheny Serviceberry ............... LC C63 La.0 T
Anmelanchier sanguinea, Rock Serviceberry ................. LC 847 T
Crataegus brainerdii, Brainerd's Hawthorn ................ LC 848 X
Crataegus phaenopyrum, Washington-thorn .................. LC 849 X
Crataegus uniflora, Dwarf Hawthorn ....................... LC 850 X
Dalibarda repens, Robin-run-away ......................... LC 210 E
Filipendula rubra, Queen-of-the-prairie .................. LC 270 P
Geum aleppicum, Yellow Avens ............................. LC 284 et.p
Geum rivale, Water Avens ................................. LC 851 0 T
Gillenia trifoliata, Bowman's-root ....................... LC 290 T
Potentilla anserina, Silverweed ...........................LC 852 1Q1 c,C,OF1L 0 P
Potentilla arguta, Tall Cinquefoil1.............~...........LC C640 E
Potentilla fruticosa, Shrubby C65 P
Potentilla palustris, Marsh Fivefinger .............~......LC 853 T
Potentilla paradoxa, Bushy Cinquefoil .................... LC 854 F- 0 T

-13-

Code No. OH
ROSACEAE. Rose Family (Cont'd.)
Prunus nigra, Canada Plum ................5...LC857 E
Prunus pumila, Great Lakes Sand Cherry . ................LC 858 X
Prunus susquehanae, Sand Cherry .............. ..LC859 E
Pyrus angustifolia, Narrow-leaved Crab ................... LC 860 X
Pyrus decora, Western Mountain-ash...... . ......LC 861 E
Rosa arkansana, Arkansas Rose ............................ LC 862 X
Rosa blandaSmooth Rose ........................LCA4l E1Q T
Rubus hispidus var. hispidus, Large Hispid Dewberry ...... LC C66 E

Rubus idaeus var. strigosus, Wild Red Raspberry ......LC 608 E P
Rubus pubescens, Dwarf Red Bramble .............LC C67 T
Rubus trivialis, Southern Dewberry ..................LC 863 X
Sanguisorba canadensis, Canadian Burniet.. ...........LC C68 P
Spiraea latifolia, Northern Meadow-sweet ..........LC 864 X
Waldsteinia fragarioides, Barren Strawberry . . ........LC C69 P

LEGUMINOSAE (FABACEAE). Pea Family
Astragalus neglectus, Cooper's Milk-vetch ................ LD 865 X
Baptisia australis, Blue False Indigo .................... LD 077 E
Baptisia leucantha, Prairie False Indigo ................. LD X59 T
Clitoria mariana, Butterfly-pea .......................... LD 866 T
Desmodium illinoense, Prairie Tick-trefoil ........... ...LD 867 E
Desmodium pauciflorum, Few-flowered Tick-trefoil ......... LD 868 E T
Desmodium sessilifolium, Sessile Tick-trefoil ............ LD 869 E
Galactia volubilis, Milk-pea ............................. LD 870 E
Lathyrus maritirnus (L. japonicus var. glaber), Inlahd

Lathyrus ochroleucus, Yellow Vetchling ................... LD 872 E
Lathyrus venosus, Wild Pea ............................... LD 873 X
Lupinus perennis, Wild Lupine ............................LD 874 P
Phaseolus polystachios, Wild Kidney Bean ................. LD 875 T
Psoralea psoralioides, False Scurf-pea ................... LD 876 T
Trifolium reflexum, Buffalo Clover ....................... LD 878 X
Trifolium stoloniferum, Running Buffalo Clover ........... LD 879 X

LINACEAE. Flax Family
Linum sulcatum, Grooved Flax ............................. LY 880 0P

OXALIDACEAE. Wood-sorrel Family
Oxalis montana, White Wood-sorrel ........................ LX 881 E

POLYGALACEAE. Milkwort Family
Polygala cruciata, Cross-leaved Milkwort ................. LZ 882 E
Polygala curtissii, Curtiss' Milkwort .................... LZ 883 E
Polygala incarnata, Pink Milkwort ......................LZ 884 E
Polygala paucifolia, Gay-wings ........................... LZ 536 E
Polygala polygama, Racemed Milkwort ...................... LZ 885 E

EUPHORBIACEAE. Spurge Family
Acalypha rhomboidea var. deamii, Deam's Three-seeded
Mercury.***************** .. .......................................... LS 886 E
Croton glandulosus, Northern Croton .................... **LS 888 T
Euphorbia polygonifolia, Seaside Spurge .................. LS 890
Euphorbia purpurea, Glade Spurge ......................... LS 269 X
Euphorbia serpens, Roundleaf Spurge ...................... LS 891 E

-14-

Code No. OH
EUPHORBIACEAE. Spurge Family (Cont'd.)
Phyllanthus caroliniensis, Carolina Leaf-flower...... ...LS 892 E

CALLITRICHACEAE. Water-starwort IFamily
Callitriche deflexa, Austin's H~ater-starviort . . ........1. 893 T

Callitriche verna (C. palustris), Water-starwort......NJ 894 AT

ANACARDIACEAE. Sumac Family
Rhus aromatica var. arenaria, Beach Sumac ..........LW 895 X
Rhus vernix, Poison Sumac ................................ LW C70 /-oL. P

AQUIFOLIACEAE. Holly Family
Ilex opaca,PAmerican Holly ................ .LQ 896 E
Nemopanthus mucronata, Catberry .......................... LQC7l P

CELASTRACEAE. Staff-tree Family
Euonyrnus americanus, American Strawberry-bush ...... ... ... LP 897 T
Pachistima canbyi, Cliff-green ........................LP 477 E

ACERACEAE. Maple Family
Acer pensylvanicum, Striped Maple. ...............LV 001 '~E

RHAMNACEAE. Buckthorn Family
Ceanothus herbaceus (C. ovatus), Prairie Redroot ......... LT 146 0E
Rhamnus alnifolia, Alder-leaved Buckthorn ................ LT 576 P
Rhamnus caroliniana, Carolina Buckthorn .................. LT 577 P

VITACEAE. Grape Family
Vitis baileyana, Possum Grape ............................ LU 898 E
Vitis cinerea, Pigeon Grape .............................. LU C72 T
Vitis labrusca, Northern Fox Grape ....................... LU C73 0 P

MALVACEAE. Mallow Family
Sida hermaphrodita, Virginia Mallow ...................... KD 899 P

HYPERICACEAE (GUTTIFERAE). St. John's Wort Family
Hypericum boreale, Northern St. John's-wort ........ ......KC 900 E
Hypericum canadense, Canadian St. John's-wort. ...... :..... KC 901 T
Hypericum denticulatum, Coppery St. John's-wort .......... KC 902 E
Hypericum ellipticum, Few-flowered St. John's-wrort ....... KC 903 X
Hypericum gymnanthum, Least St. John's-wort .............. KC 904 E
Hypericum hypericoides (Ascyrum hypericoides), St.
Andrew's Cross................ .... ........ .... ...... .KC C74 P
Hypericum kalmianum, Kalm's St. John's-wort .............. KC 905 T
Hypericum majus, Tall St. John's-wort .................... KC 906 El o T
Hypericum tubulosum (including H. walteri), Large Marsh
St. John' s-wort. ... .. .. .. . ... .. .. . ... .. .. . .. .. .. .. ... KC 907 T

CISTACEAE. Rockrose Family
Helianthemum bicknellii, Plains Frostweed ................ KK 309 ET
Helianthemum canadense, Canada Frostweed ................. KK 909 T
Hudsonia tomentosa, Beach-heather ....................KK 339 X

Code No. OH
CISTACEAE.- Rockrose Family (Cont'd.)
Lechea intermedia, Round-fruited Pinweed . . ........KK A45 T
Lechea leggettii, Leggett's Pinweed ............KK 910 T
Lechea minor, Thyme-leaf Pn inw............... .KK 911 E
Lechea stricta, Prairie Pinweed . ..............KK 912 X
Lechea tenuifolia, Narrow-leaved Pinweed ..........KK A46 E
Lechea villosa, Hairy Pin wed............... . .KK C75 P

VIOLACEAE. Violet Family
Viola hirsutula, Southern Wood Violet..............KH C76 P
Viola lanceolata, Lance-leaved Violet ....... . ...KH 913 E
Viola missouriensis, Missouri Violet ...............KH 914 X
Viola nephrophylla, Northern Bog Violet ..........KH 915 Ejo T
Viola pallens, Northern White Violet ...............KH C77 P
Viola pedata, Bird-foot Violet ........................... KH 753 T
Viola pedatifida, Prairie Violet ................ . .....KH916 X
Viola primulifolia, Primrose-leaved Violet ...........KH 917 E
Viola rotundifolia, Round-leaved Violet ...........KH 918 P
Viola tripartita var. glaberrima, Wedge-leaf Violet ...... KH 754 T
Viola tripartita var. tripartita, Three-parted Violet .... KH 754 E
Viola walteri, Walter's Violet .................. . .....KH 920 T

PASSIFLORACEAE. Passion-flower Family
Passiflora incarnata, Passion-flower ..................... KJ 921 T

CACTACEAE. Cactus Family
Opuntia compressa (0. humifusa), Prickly Pear ............ JM C78 P

ELAEAGNACEAE. Oleaster Family
Shepherdia canadensis, Canadian Buffalo-berry ............ LL 922 T

MELASTOMATACEAE. Melastoma Family
Rhexia virginica, Virginia Meadow-beauty ................. LK 581 T

ONAGRACEAE. Evening-primrose Family
Epilobium angustifolium, Fireweed ...... .................LJ E99 T
Epilobium strictum, Simple Willow-herb ................... LJ 923 T
Oenothera pilosella, Meadow Sundrops ..................... LJ 925 P
Oenothera triloba, Sessile Evening-primrose .............. LJ 924 X

HALORAGACEAE. Water-mil1foil Family
Myriophyllum exalbescens, American Water-milfoil ......... LF 926 iL2jO P
Myriophyllum heterophyllum, Two-leaved Water-milfoil ..... LF 927 E
Myriophyllum verticillatum, Green Water-milfoil .......... LF 928 X

ARALIACEAE. Ginseng Family
Aralia hispida, Bristly Sarsaparilla ..................... L2 929 X
Aralia spinosa, Devil's-walkingstick ..................... L2 C79 P
Panax quinquefolius, American Ginseng .................... L2 480 Lo~P

UMBELLIFERAE (APIACEAE). Parsley Family
Chaerophyllum procumibens var. shortii, Short's Chervil ...L3 930 P
Eryngium yuccifolium, Rattlesnake-master ................. L3 932 T

-1 6-

Code No. OH
UMBELLIFERAE (APIACEAE). Parsley Family (Cont'd.)
Hydrocotyle americana, American Water-pennywort ......L3 343 P
Hydrocotyle umbel lata, Navelviort....... . ......L3 933 E
Ligusticum canadense, American Lovage ...........L3 934 E
Perideridia americana, Perideridia...... . ......L3 503 E
Thaspium pinnatifidum, Cut-leaf Meadow-parsnip.......L3 718 T

CORNACEAE. Dogwood Family
Cornus canadensis, Bunchberry ...... . ........LM 935 T
Cornus rugosa, Round-leaved Dogwood ............LM A94 A.'0 T

PYROLACEAE. Pyrola Family
Chimaphila umbellata, Pipsissewa ...... . .......KQ 153 T
Pyrola secunda, One-sided Wintergreen ....... . ...KQ 936 X

ERICACEAE. Heath Family
Andromeda glaucophylla, Bog-rosemary .............KQ 030 X
Arctostaphylos uva-ursi, Bearberry .............. ........KQ 937 X
Chamaedaphne calyculata, Leather-leaf ............KQ 938 P
Gaultheria hispidula, Creeping Snowberry ..............KQ 939 X
Ledum groenlandicum, Labrador-tea .....................KQ 940 E
Lyonia ligustrina, Maleberry ............................. KQ 941 X
Rhododendron calendulaceum, Flame Azalea..........KQ 942 E
Rhododendron maximum, Great Rhododendron ............ .KQ 943 T
Rhododendron nudiflorum, Pinxter-flower . ...............KQ 944 T
Rhododendron roseum, Northern Rose Azalea .............KQ 588 T
Vaccinium atrococcum, Black Highbush Blueberry ........... KQ C8O T
Vaccinium caesium, Small Deerberry .................. ...KQ 945 E
Vaccinium macrocarpon, Large Cranberry ................... KQ 746 P
Vaccinium myrtilloides, Velvet-leaf Blueberry ............ KQ 946 X
Vaccinium oxycoccos, Small Cranberry .... ...............KQ 747 E

PRIMULACEAE. Primrose Family
Dodecatheon meadia, Shooting-star . .............KW 225 P
Hottonia inflata, Featherfoil ............................ KW 334 E
Trientalis borealis, Star-flower ...................KW 730 P

STYRACACEAE. Storax Family
Halesia carolina, Silverbell....... . ........KU 304 E
Styrax americana, Snowbell .................KU 948 E
Styrax grandifolia, Bigleaf Snowbell ..................... KU 949 X

OLEACEAE. Olive Family
Chionanthus virginicus, Fringe-tree ...................... NN 950 E
Fraxinus tomentosa, Pumpkin Ash ................ ........NN 951 X

GENTIANACEAE. Gentian Family
Bartonia virginica, Yellow Bartonia ...................... NB C81 P
Gentiana alba (G. flavida), Yellowish Gentian ............ NB 281 E
Gentiana clausa, Closed Gentian . .......................NB 952 4 T
Gentiana crinita, Fringed Gentian ........................ NB 283 .4 P
Gentiana procera, Small Fringed Gentian .................. NB C82 P
Gentiana puberulenta (G. puberula), Prairie Gentian ...... NB 953 E
Gentiana quinquefolia, Stiff Gentian.................. .*..NB X60 C. P

"17-

Code No. OH
GENTIANACEAE. Gentian Family (Cont'd.)
Gentiana saponaria, Soapwort Gentian ................NB 954 E
Gentiana villosa, Sampson's Snakeroot .................... NB 955 E
Obolaria virginica, Pennywort ............................ NB C83 P
Swertia caroliniensis, American Columbo .................. NB 705 P

MENYANTHACEAE (GENTIANACEAE). Buckbean Family
Menyanthes trifoliata, Buckbean ...................,.......NB 448 E

APOCYNACEAE. Dogbane Family
Apocynum sibiricurm (incl. var. cordigerum), Clasping-
leaf Dogbane .. ..... .................. ... ~ .........NC 956 ,9EO Icc P

ASCLEPIADACEAE. Milkweed Family
Asclepias amplexicaulis, Bluntleaf Milkweed ............ND C84 P
*Asclepias variegata, White Milkweed ...................... ND C85 T
Asclepias viridis (Asclepiodora viridis), Spider
Mil1kweed. . ... . ..... .. .. ... .. .. ... . .. .. . .. .. . ... ... ND 957 T
Matalea obliqua (Gonolobus obliquus), Angle-pod .......... ND C24 P

CONVOLVULACEAE. Morning-glory Family
Cuscuta compacta, Sessile Dodder ......................... NE 958 1
Cuscuta coryli, Hazel Dodder ............................. NE 959 0 T
Cuscuta glomerata, Glomerate Dodder ...................... NE 960 X
Cuscuta pentagona, Five-angled Dodder .................... NE 961 T

POLEMONIACEAE. Phlox Family
Phlox glaberrima, Smooth Phlox ........................... NF 962 P
Phlox ovata, Mountain Phlox .............................. NF 963 T
Phlox stolonifera, Creeping Phlox . .............NF 964 P
Polemonium reptans var. villosum, Braun's Jacob's-ladder.NF 531 P

HYDROPHYLLACEAE. Waterleaf Family
Phacelia bipinnatifida, Fern-leaf Scorpion-weed .......... NG 965 T
Phacelia dubia, Small-flowered Scorpion-weed ............. NG 508 X
Phacelia ranunculacea, Blue Scorpion-weed ................ NG 510 E.

BORAGINACEAE. Borage Family
Cynoglossum boreale, Northern Wild Comfrey ............... NH 966 E
Lithospermum cr-oceuri, Plains Puccoon ..................... NH DLO P
Myosotis macrosperma, Bristly Scorpion-grass ............. NH 968 T
Onosmodium hispidissimum, False Gromwell ................. NH 969 p

LABIATAE (LAMIACEAE). Mint Family
Collinsonia verticillata (Micheliella verticillata),
Early Stoneroot ....... *.. ............. ..........NL 170 E
Hedeoma hispida, Rough Pennyroyal ........................ NL 970 E
Meehania cordata, Meehania ............................... NL 444 P
Monarda punctata, Dotted Horsemint ....................... NL 971 C~E
Pycnanthemum muticum, Blunt Mountain-mint ................ NL A96 T
Pycnanthemum pilosum, Hoary Mountain-mint ................ NL 972 E

Satureja arkansana, Limestone Savory ..................... NL C88 EtP

*Asclepias viridiflora, Green Milkweed .................... ND DK7O P

Code No. OH
LABIATAE (LAMIACEAE). Mint Family (Cont'd.)
Scutellaria integrifolia, Hyssop Skullcap .........NL C89 P
Scutellaria saxatilis, Rock Skullcap...... . .....NL 651 E
Scutellaria serrata, Showy Skullcap..............NL C9O P
Synandra hispidula, Synandra...... . .........NL 708 T
Trichostema dichotomum var. lineare (T. setaceum),
Narrow-leaved Bluecurls .. . ....... ..........NL 974 E

SCROPHULARIACEAE. Figwort Family.
Agalinis gattingeri (Gerardia gattingeri), Gattinger's
Foxglove ........... ... ............. ........NP 975 X
Agalinis paupercula var. paupercula (Gerardia paupercula
var. paupercula), Small Purple Foxglove ...........NP 976 E
Agalinis paupercula var. borealis (Gerardia paupercula
var. borealis), Northern Purple Foxglove ............ NP 976 X
Agalinis skinneriana (Gerardia skinneriana), Skinner's
Foxglove .. .....................................NP 977 E
Aureolaria pedicularia var. pedicularia (Gerardia
pedicularia var. pedicularia), Woodland Fern-leaf
False Foxglove............... . .. ... ......NP 978 X
Aureolaria pedicularia var. ambigens (Gerardia
pedicUlaria var. ambigens), Prairie Fern-leaF
False Foxgl ove. .. ........ . .. . .. .. .. .. . .... .. . ... ..NP 978 E
Besseya bullil (Wulfenia bullii), Besseya ................ NP 979 X
Buchnera americana, Bluehearts ........................... NP 980 T
Gratiola virginiana, Round-fruited Hedge-hyssop .......... NP 981 T
Gratiola viscidula var. shortii, Short's Hedge-hyssop .... NP 982 T
Linaria canadensis, Old-field Toadflax ................... NP 983 E
Melampyrum lineare, Cow-wheat ............................ NP 984 T
Penstemon canescens, Gray Beard-tongue ................... NP 986 E
Penstemon laevigatus, Smooth Beard-tongue .........NP 987 T
Penstemon pallidus, Downy White Beard-tongue ............. NP C91 T
Penstemon tubaeflorus, White-wand Beard-tongue .... ... .... NP 988 X
Tomanthera auriculata (Gerardia auriculata), Ear-leaf
Foxglove.. .. . . NP 722 E
Veronica angli-qaia ro-ipre.......PC92 P

BIGNONIACEAE. Bignonia Family
Bignonia capreolata, Cross-vine .......................... NZ C93 P

OROBANCHACEAE. Broomrape Family
Orobanche ludoviciana, Louisiana Broom-rape .............. NQ 989 E

LENTIBULARIACEAE. Bl adderwort Family
LUtricularia cornuta, Horned Bladderwort ................... NS 990 E
Utricularia gibba, Humped Bladderwort .................... NS A59 T
Utricularia intermedia, Flat-leaved Bladderwort .......... NS 991 T
Utricularia minor, Lesser Bladderwort .....................NS 992 T

ACANTHACEAE. Acanthus Family
Ruellia caroliniensis, Carolina Ruellia .................. NR 993 p
Ruellia humilis, Low Ruellia ............................. NR 61l P

PLANTAGINACEAE. Plantain Family
Plantago cordata, Heart-leaf Plantain .....................NM .527 E

Code No. OH
RUBIACEAE. Madder Family
Galium labradoricum, Bog Bedstraw..............NU 995 E
Galium palustre, Marsh Bedstraw ......... . .....NU 996 E
Houstonia nigricans, Narrow-leaved Summer Bluets......NU C94 E1O T
Houstonia purpurea, Large Summer Bluets...........Nu C95 P
Spermacoce glabra, Smooth Buttonweed ........ . ...NU C96 P

CAPRIFOLIACEAE. Honeysuckle Family
Linnaea borealis, American Twinflower . . . ...... . ..NV 402 X
Lonicera canadensis, Canada 'Fly-honeysuckle . . . ......NV 411 p
Lonicera flavida, Pale Yellow Honeysuckle . . . .......NV 997 X
Lonicera oblongifolia, Swamp Fly-honeysuckle . . . ......NV 998 X
Lonicera prolifera, Grape Honeysuckle ......... . ..NV 999 T
Lonicera villosa, Mountain Fly-honeysuckle. . . .......NV A0l X
Symphoricarpos albus var. albus, Snowberry . . . .......NV A02 0 E
Viburnum alnifolium, Hobblebush .................NV A03 P
Viburnum molle, Soft-leaved Arrow-wood . . . ...... . ..NV A04 E
Viburnum opulus var. americanum (V. trilobum),
H ighbush-cranberry. . ... .. . ... .. . . ... .. .. . .. . .....NV B07 P
Viburnum rufidulum, Southern Black-haw . . . ...... . ..NV A05 E

VALERIANACEAE. Valerian Family
Valeriana ciliata, Prairie Valerian .......... . ...NW A06 T
Valeriana uli giniosa, Swamp Valerian . ............NW X23 X
CAMPANULACEAE. Bell flower Family
Campanula rotundifolia, Harebell..............NT B02 EJ0T

COMPOSITAE (ASTERACEAE). Composite Family
Artemisia caudata, Beach Wormwood ........................ NX K07 EOARC.. T
Aster acuminatus, Mountain Aster ......................... NX KO8 X
Aster dumosus, Bushy Aster ...................NX K09 CIO T
Aster infirmus, Weak Aster ..........................NX Kll T
Aster linariifolius, Stiff-leaf Aster .................... NX 069 p
Aster oblongifolius, Shale Barren Aster...............NX K12 T
Aster ptarmiicoides, Yarrow Aster .........................NX K13 X
Aster solidagineus (Sericocarpus linifolius), Narrow-
leaved Aster ... .. ................ . ......NX C14 T
Aster surculosus, Creeping Aster ......................... NX K15 X
Cacalia muhlenbergii, Great Indian-plantain .............. NX K17 T
Cacalia tuberosa, Fen Indian-plantain .................... NX E02 T
Chrysogonum virginianum, Golden-knees .................... NX 155 p
Cirsium carolinianum, Carolina Thistle ................... NX K18 E
Cirsium hillii, Hill's Pasture Thistle ................... NX K19 T

Echinacea purpurea, Purple Coneflower .................... NX 239 P
Erigeron divaricatus, Bushy Horseweed .................... NX K21 E
Eupatorium album, White Thoroughwort ..................... NX K22 E
Eupatorium aromaticum, Small White Snakeroot ............. NX K23 E
Eupatorium hyssopifolium, Hyssop Thoroughwort ............ NX K24 E
Eupatorium incarnatum, Pink Thoroughwort ................. NX K25
Eupatorium rotundifolium (including E. pubescens),
Roundleaf Thoroughwort .......... . .......NX K26 P
Gnaphaliumn viscosum (G. macounii), Winged Cudweed ........ NX K27 X
*Diodia virginiana, Southern Buttonweed ................... NU DOl P

-20-
Code No. OH
COMPOSITAE (ASTERACEAE). Composite Family (Cont'd.)
Helianthus mollis, Ashy Sunflower.. . ............NXA63 T
Helianthus occidentalis, Western Sunflower ............... NX K28 T
Heterotheca mariana (Chrysopsis mariana), Golden-aster ...0NX K(30 P
Heterotheca graminifolia (Chrysopsis graminifolia),
Silkgrass.................... ...........NX K(29 E
Hieracium canadense, Canada Hawkweed ....... ............NX K(31 E
Hieracium marianum, Maryland Hawkweed .............NX K32 X
Hymenoxys acaulis (Actinea herbacea), Lakeside Daisy. ...NX K(330 E
Krigia dandelion, Potato-dandelion .......................NX K(34 E
Krigia virginica, Dwarf Dandelion . ................NX K(35 T
Kuhnia. eupatorioides (including varieties; Brickellia
eupatorioides), False Boneset ................... ...NXK16 C-P
Liatris cylindracea, Slender Blazing-star ................ NX EIO T
Liatris novae-angliae (L. novae-angliae var. nieuwlandii
=X L. nieuwlandii in Fernald 1950), New England

Liatris scabra, Large Blazing-star ....................... NX K(36 E
Liatris squarrosa, Scaly Blazing-star.. . ...........NX K(37 T
Megalodonta beckii, Water-marigold . ........ . ...NX K38 X
Pluchea camphorata, Camphorweed .......................... NX A65 X
Prenanthes aspera, Rough Rattlesnake-root ................ NX K(39 X
Prenanthes crepidinea, Nodding Rattlesnake-root .......... NX K(40 T
Prenanthes racemosa, Prairie Rattlesnake-root ............ NX 1(41 T
Senecio pauperculus, Balsam Squaw-weed...........NX 659 9:10 p
Silphium laciniatum, Compass-plant ....................... NX E13 E
Solidago arguta, Cut-leaf Goldenrod ...................... NX 1(42 E
Solidago odora, Sweet Goldenrod .......................... NX K(43
Solidago ohioensis, Ohio Goldenrod ....................... NX K44 P
Solidago remota, Great Lakes Goldenrod ................... NX K45 T
Sol idago ri ddelIIi i ,Ri ddel 1 1s Gol denrod ................... NX K46 0P
Solidago rigida, Stiff Goldenrod .........................NX K(47 EQP
Solidago squarrosa, Leafy Goldenrod ...................... NX 683 T
Verbesina helianthoides, Hairy Wing-stem .................NX 1(48 P
Verbesina occidentalis, Yellow Crownbeard ................ NX K(49 E
Vernonia fasciculata, Prairie Ironweed ................... NX 1(50 E

APPENDIX C

COUNTY LISTS OF RARE PLANTS OF THE COASTAL ZONE

ASHTABULA COUNTY

Ohio Status Scientific Name Common Name

E Acer penszivanic-zc m Striped Maple
T AmmophiZa breviZiguZata American Beach Grass
P Apocynum sibiricum Clasping-leaf Dogbane
T Artemisia caudata Beach Wormwood
P Cakile edentuZa var. Zacustris Inland Sea-rocket
T CaZZitriche verna Water-starwort
E Carex aquatiZis Leafy Tussock Sedge
T Carex aurea Golden-fruited Sedge
T CZaytonia caroZiniana Carolina Spring-beauty
T Cornus rugosa Round-leaved Dogwood
P Cyperus schweinitzii Schweinitz's Umbrella-sedge
P Euphorbia poZygonifoZlia Seaside Spurge
T Gentiana clausa Closed Gentian
P Gentiana crinita Fringed Gentian
T Juncus aZpinus Alpine Rush
T Juncus baZticus Baltic Rush
T Lathyrus maritimus Inland Beach-pea
P Liparis Zoeselii Fen Orchid
P MyriophyZZllum exalbescens American Water-milfoil
P Nlajas 'Zexilis Slender Naiad
P Potamogeton epihydrus Nuttall's Pondweed
T Potamogeton richardsonii Richardson's Pondweed
P PotentiZZla anserina Silverweed
P Sporobolus cryptandrus Sand Dropseed
P Triplasis purpurea Purple Sand-grass
P VaZZlisneria americana Eel-grass

LAKE COUNTY

Ohio Status Scientific Name Common Name

T Amnelanchier Zaevis Allegheny Serviceberry
T Ammophila breviZiguZata American Beach Grass
P Arabis lyrata Lyre-leaf Rock-cress
P CakiZe edentuZa var. Zacustris Inland Sea-rocket
E Carex aquatilis Leafy Tussock Sedge
P Euphorbia poZygonifoZia Seaside Spurge
T Lathyrus martimus Inland Beach-pea
P Myriophyllum exalbescens American Water-milfoil
P Panax quinquefolius American Ginseng
T Potamogeton natans Floating Pondweed
P PotentilZa anserina Silverweed
P Rhus vernix Poison Sumac
P SporoboZus cryptandrus Sand Dropseed
P Triplasis purpurea Purple Sand-grass
P VallZZisneria americana Eel-grass

CUYAHOGA COUNTY

T AmmophiZa breviZiguZata American Beach Grass
T Artemisia caudata Beach Wormwood
P CakiZe edentula var. lacustris Inland Sea-rocket
P Cyperus schweinitzii Schweinitz's Umbrella-sedge
P Euphorbia polygonifoZia Seaside Spurge
P Gentiana quinquefolia Stiff Gentian
P Kuhnia eupatorioides False Boneset
T Lathyrus maritimus Inland Beach-pea
T LeptoZoma cognatwn Fall Witch-grass
E Monarda punctata Dotted Horsemint
P PotentiZZa anserina Silverweed
P SporoboZus cryptandrus Sand Dropseed
P Triplasis purpurea Purple Sand-grass
P VaZZisneria americana Eel-grass

LORAIN COUNTY

P CakiZe edentuZa var. Zacustris Inland Sea-rocket
P Euphorbia poZygonifoZia Seaside Spurge
P Triplasis purpurea Purple Sand-grass

ERIE COUNTY

Ohio Status Scientific Name Common Name

T AmmophiZa breviZiguZata American Beach Grass
P Apoeynwum sibiricum Clasping-leaf Dogbane
T Artemisia caudata Beach Wormwood
T Aster dumosus Bushy Aster
P Betula popuZifoZia Gray Birch
P CakiZe edentuZa var. Zacustris Inland Sea-rocket
T Campanula rotundifoZia Harebell
P Carex eburnea Bristle-leaf Sedge
T Carex lasiocarpa Slender Sedge
P Carex trichocarpa Hairy-fruited Sedge
T Carex viridula Little Green Sedge
T Cyperus diandrus Low Umbrella-sedge
P Cyperus engelmannii Engelmann's Umbrella-sedge
P Cyperus schweinitzii Schweinitz's Umbrella-sedge
T Desmodium pauciflorum Few-flowered Tick-trefoil
P EZeocharis compressa Flat-stem Spikerush
T EZeocharis eZliptica Yellow-seeded Spikerush
P Euphorbia poZygonifoZia Seaside Spurge
P Geum aleppicum Yellow Avens
T Helianthemum bickneZZii Plains Frostweed
T Houstonia nigricans Narrow-leaved Summer Bluets
T Hypericum majus Tall St. John's-wort
E Iris brevicaulis Leafy Blue Flag
T Juncus alpinus Alpine Rush
T Juncus scirpoides False Bulrush
T Juncus secundus One-sided Rush
P Liparis ZoeseZii Fen Orchid
P Lophotocarpus calycinus Southern Wapato
P MyriophyZZum exalbescens American Water-milfoil
P Najas flexilis Slender Naiad
T PoZygonum pensyZvanicumvar. egZanduZcsum Lake Erie Pinkweed
P PotentiZZa anserina Silverweed
T PotentilZa paradoxa Bushy Cinquefoil
T Ranunculus fascicuZaris Early Buttercup
T Rosa blanda Smooth Rose
P Rubus idaeus var. strigosus Wild Red Raspberry
T Sagittaria cuneata Wapato
P Satureja arkansana Limestone Savory
T ScZeria verticiZZata Low Nut-rush
P Senecio pauperculus Balsam Squaw-weed
T Sisyrinchium montanum Northern Blue-eyed-grass
T Sisyrinchium mucronatum Narrow-leaved Blue-eyed-grass
P SoZidago rigida Stiff Goldenrod
T Spiranthes magnicamporum Great Plains Ladies'-tresses
P SporoboZus cryptandrus Sand Dropseed
P Triplasis purpurea Purple Sand-grass
P VaZZisneria americana Eel-grass
T Viola nephrophylla Northern Bog Violet

OTTAWA COUNTY

Ohio Status Scientific Name Common Name

P Agropyron trachycaulum Bearded Wheat Grass
T Ammophila breviZigulata American Beach Grass
T Anemone cyZindrica Prairie Thimbleweed
P Apocynum sibiricum Clasping-leaf Dogbane
E Arabis drwmmondii Drummond's Rock-cress
T Arabis hirsuta var. pycnocarpa Western Hairy Rock-cress
T Arenaria stricta Rock Sandwort
T Artemisia caudata Beach Wormwood
P AscZepias viridiflora Green Milkweed
T Aster dumosus Bushy Aster
P Cakile edentuZa var. Zacustris Inland Sea-rocket
T CampanuZa rotundifoZia Harebell
T Carex atherodes Wheat Sedge
E Carex aurea Golden-fruited Sedge
P Carex brevior Tufted Fescue Sedge
T Carex crawei Crawe's Sedge
P Carex eburnea Bristle-leaf Sedge
P Carex ZupuZliformis False Hop Sedge
P Carex muhlenbergii Muhlenberg's Sedge
E Carex sprengeZii Sprengel's Sedge
T Carex viridula Little Green Sedge
E Ceanothus herbaceus Prairie Redroot
P Celtis tenuifolia Dwarf Hackberry
T CeratophylZZum echinatum Prickly Hornwort
E Chenopodiwn leptophyZZlwnum Slender Goosefoot
T CZadium mariscoides Twig-rush
T Cornus rugosa Round-leaved Dogwood
T CorydaZis sempervirens Rock-harlequin
T Cuscuta coryli Hazel Dodder
T Cyperus diandrus Low Umbrella-sedge
P Cyperus engelmannii Engelmann's Umbrella-sedge
P Cyperus schweinitzii Schweinitz's Umbrella-sedge
P Descurainia pinnata Tansy-mustard
P EZeocharis compressa Flat-stem Spikerush
T EZeocharis intermedia Matted Spikerush
T EZeocharis olivacea Olivaceous Spikerush
P Equisetum variegatum Variegated Scouring-rush
P Euphorbia poZygonifoZia Seaside Spurge
T Geum rivaZe Water Avens
T Houstonia nigricans Narrow-leaved Summer Bluets
E Hymenoxys acaulis Lakeside Daisy
T Hypericum majus Tall St. John's-wort
T Juncus alpinus Alpine Rush
T Juncus balticus Baltic Rush
E Juncus interior Interior Rush
P Lilium philadeZphicum Wood-lily
P Linum suZlcatum Grooved Flax
P Liparis loeseZii Fen Orchid
P Lophotocarpus calycinus Southern Wapato
P MyriophyZZllum exalbescens American Water-milfoil
P Najas fZexiZis Slender Naiad
E Nuphar variegatum Bullhead Lily

OTTAWA COUNTY (Continued)

Ohio Status Scientific Name Common Name

X PZatanthera Zeucophaea Prairie Fringed Orchid
T PoZygonum pensyZvanicwum var. eglanduZosum Lake Erie Pinkweed
E Polygonum ramosissimum Bushy Knotweed
E PopuZus balsamifera Balsam Poplar
E Potamogeton filiformis Filiform Pondweed
T Potamogeton natans Floating Pondweed
T Potamogeton richardsonii Richardson's Pondweed
T Potamogeton zosteriformis Flat-stem Pondweed
P PotentiZZa anserina Silverweed
E PotentiZZa arguta Tall Cinquefoil
T PotentiZZla paradoxa Bushy Cinquefoil
T RanuncuZus fascicularis Early Buttercup
T Rosa blanda Smooth Rose
T Sagittaria cuneata Wapato
T Sagittaria graminea Grass-leaf Arrowhead
P Salix bebbiana Long-beaked Willow
P Satureja arkansana Limestone Savory
T Scirpus purshianus Pursh's Bulrush
E Scirpus smithii Smith's Bulrush
P Senecio pauperculus Balsam Squaw-weed
T Sisyrinchium mucronatum Narrow-leaved Blue-eyed-grass
P Solidago riddellii Riddell's Goldenrod
P SoZlidago rigida Stiff Goldenrod
T Spiranthes magniccamporumw Great Plains Ladies'-tresses
P Sporobolus cryptandrus Sand Dropseed
E Symphoricarpus aZbus var. albus Snowberry
P Thuja occidentalis Arbor Vitae
P Triplasis purpurea Purple Sand-grass
P VaZZllisneria americana Eel-grass
T VioZa nephrophyZZlla Northern Bog Violet
P Vitis Zlabrusca Northern Fox Grape
T Zizania aquatica Wild Rice

LUCAS COUNTY

Ohio Status Scientific Name Common Name

T AmmophiZa breviZiguZata American Beach Grass
T Anemone cyZindrica Prairie Thimbleweed
P Apocynum sibiricum Clasping-leaf Dogbane
E Arabis divaricarpa Limestone Rock-cress
P CakiZe edentula var. Zacustris Inland Sea-rocket
T Cyperus diandrus Low Umbrella-sedge
P Cyperus engeZmannii Engelmann's Umbrella-sedge
P Cyperus schweinitzii Schweinitz's Umbrella-sedge
P EZeocharis compressa Flat-stem Spikerush
T Eleocharis intermedia flatted Spikerush
P Euphorbia poZygonifoZia Seaside Spurge
P Geum aleppicum Yellow Avens
P Lophotocarpus calycinus Southern Wapato
E MuhZenbergia cuspidata Plains Muhlenbergia
E Nuphar variegatum Bullhead Lily
X PZatanthera Zeucophaea Prairie Fringed Orchid
T Potamogeton natans Floating Pondweed
P PotentilZZa anserina Silverweed
T PotentiZZa paradoxa Bushy Cinquefoil
T Sagittaria cuneata Wapato
P SaZix bebbiana Long-beaked Willow
T Sparganium androcladum Keeled Bur-reed
P SporoboZus cryptandrus Sand Dropseed
P TripZasis purpurea Purple Sand-grass
P VaZZisneria americana Eel-grass

SANDUSKY COUNTY

P Descurainia pinnata Tansy-mustard
E Juncus interior Inland Rush

WOOD COUNTY

None

APPENDIX D

OHIO PLANTS OF FEDERAL CONCERN

1/28/81

OHIO PLANTS OF FEDERAL CONCERN

Summarized by the
Division of Natural Areas and Preserves
Ohio Department of Natural Resources
Fountain Square, Bldg. F
Columbus, Ohio 43224

KEY

Federal Status: (From Federal Register, Vol. 45, No. 242, December 15, 1980)

T = Species currently listed as threatened by U.S. Fish & Wildlife Service

I = F&WS presently has sufficient information on hand to support the
biological appropriateness of taxa being listed as endangered or
threatened species. Development of proposed and final rules may
require several years.

2 = F&WS currently has information which indicates the probable approp-
riateness of listing as endangered or threatened, but sufficient
information is not presently available to biologically support a
proposed rule.. Further biological research will usually be necessary
to determine status of the taxa.

STATE STATUS

E = State of Ohio Endangered Species, from criteria in Ohio Administrative
Rule 1501:18-1-02, effective July 14, 1980.
T = State of Ohio Threatened Species, from criteria in Ohio Administrative
Rule 1501:18-1-02, effective July 14, 1980.

X = A native Ohio plant species presumed extirpated from the state.

P = A native Ohio plant species designated as potentially threatened,
from criteria listed in "Rare Species of Native Ohio Wild Plants",
DNAP, March, 1980.

1/28/81

Federal State
Status Status Scientific Name Common Name(s)

T E Aconitum noveboracense Northern Wild Monkshood

2 X Besseya buZZii Besseya

1 E Calamagrostis insperata Ofer Hollow Reed Grass
Bartley's Reed Bent Grass

1 E Hymenoxys acauZis var. gZabra Lakeside Daisy

2 E Paxistima canbyi Cliff-green

1 E PZantago cordata Heart-leaved Plantain
Heart-leaf Plantain

1 X* PZatanthera leucophaea Prairie White-fringed Orchid
Prairie Fringed Orchid

2 X Poa paZudigena Blue Grass
Marsh Spear-grass

2 P PoZemonium reptans var. viZZoswn Jacob's Ladder
Braun's Jacob's-ladder

2 X Potamogeton hZZiii Hill's Pondweed

2 X** Rhus triZobata var. arenaria Sumac
(Synonym Rhus aromatica var. arenaria) Beach Sumac

2 E SiZene regia Royal Catchfly

1 P SuZZivantia suZZivantii Sullivantia

2 T Synandra hispiduZa Synandra

1 E TroZZlius Zaxus ssp. Zaxus Spreading Globe-flower

2 X Valeriana uliginosa Swamp Valerian
* Ohio populationsdiscovered in 1980. Official Ohio list will be amended in 1981 to
reclassify species to endangered status.

** Possible Ohio populations discovered, which will be verified by June 1981.

APPENDIX E

ADMINISTRATIVE RULES FOR O.R.C. CHAPTER 1518

Adopted: July I,' 1980
1501:18-2-04. Legally Obtained Plants. Any nurseryman, dealer, or individual Effective: July 14, 1980
who has legally obtained any native species of wild plants listed as endangered or
threatened in Rules 1501:18-1-03 and 1501:18-1-04 of the Administrative Code from
another state or commercial propagator shall provide proof upon demand that Administrative Rules
said plants were legally acquired. for

1501:18-2-05. Exceptions. Nothing herein shall be interpreted in such a manner Ohio Revised Code Chapter 1518
as to prevent any nurseryman or dealer who is licensed under Chapter 927. of the
Revised Code from selling, offering for sale, shipping, or otherwise disposing of "Ohio Endangered Plant Law'
any endangered or threatened species of plants or parts thereof when such
plants have been commercially grown by a licensed nursery or legally imported
into this state. Furthermore, nothing herein shall prohibit a person from willfully
rooting up, injuring, destroying, removing, taking, or possessing an endangered
or threatened plant from his property or from the property became effective on August 23, 1978 after passage of Amended
person has written permission of the owner, lessee, or other person entitled to Substitute House Bill No. 908 on April 25, 1978. Rules 1501:18-1-03
possession. and 1501:18-1-04 identify the species designated as endangered and
threatened and are available separately from this material. The
species lists may be requested from the Division of Natural Areas
and Preserves, Ohio Department of Natural Resources, Fountain
Square, Columbus, Ohio 43224.

1501:18-1-01. Definitions. As used in rules 1501:18-1-01 to 1501:18-2-05 of the
Administrative Code:
(A) "Commercial Purposes" means with intent to sell or trade for gain or profit.
(1) "Commercially Grown" means grown under cultivation in tilled plots or in a
greenhouse.
(C) "Division" means the Ohio Department of Natural Resources, Division of
Natural Areas and Preserves.
(D) "Endangered Species" means a native Ohio plant species whose natural
populationls) are in immediate danger of extirpation from Ohio, and/or any
federal endangered plant species in Ohio.
(E) "Chief" means Chief of the Division of Natural Areas and Preserves.
(F) "Federal Endangered Species" means all species native to this state which
are listed as endangered on the United States list of endangered and
threatened wildlife and plants pursuant to the Endangered Species Act of
1973, 87 Stat. 884, 16 U.S.C. 1531, as amended.
(G) "Federal Threatened Species" means all species native to this state which
are listed as threatened on the United States list of endangered and
threatened wildlife and plants pursuant to the Endangered Species Act of
1973, 87 Stat. 884, 16 U.S.C. 1531, as amended.
(H) "Native Habitat" means the environment in which a species could exist as a
natural population.
(I) "Native Species" means a species which, by accepted scientific evidence,
was present in Ohio immediately prior to European exploration and
settlement.

(J) "'Natural Population" means one or more plants of a single species which,
by lack of contrary evidence, were apparently established in a given area 1501:18-1-04. Threatened Species. The following native species of plants are
through dispersal by natural means, excluding dispersal by humans, human designated as threatened with becoming endangered in the state:
transportation systems, or planting by humans. (see species list)
(K) "Person" means any individual, firm, partnership, corporation, company,
association, or any combination of individuals, or any employee, agent, or 1501:18-2-01. Commercial Taking of Plants. It shall be unlawful for any
officer thereof. person to take native species of wild plants or parts thereof that are listed as
(L) "Plant" means any member of the plant kingdom, including but not limited endangered by rule 1501:18-1-03 of the Administrative Code, or threatened by rule
to: roots; stems, including bulbs, corms, tubers, rhizomes, stolons, and other 1501:18-1-04 of the Administrative Code, for commercial purposes from any
stem forms; leaves; flowers; fruits; cones; seeds; spores; and gametophytes. woodlot, field, or forest, or from any other location in which such plant is found
(M) "Species" means a species, subspecies or variety. growing in its native habitat.
(N) "Threatened Species" means a native Ohio plant species likely to become an
endangered species in Ohio within the foreseeable future, and/or any
federal thngereatened plant species in Ohio winot on thforeseeable fustate end/or angered 1501:18-2-02. Collecting Endangered and Threatened Plants. It shall be
federal threatened plant species liin Ohio not on the state endangered unlawful for any person to take, possess or transport for botanical, educational
(O) "Wild Plant" means an individual member of a natural plant population. and scientific purposes, or for propagation in captivity to preserve the species,
any native Ohio species of wild plants or parts thereof, that are listed as
endangered by rule 1501:18-1-03 of the Administrative Code or threatened by rule
identifying and1501:18-1-02. Designation riteria. The following criteria shall bare inused fodanger 1501:18-1-04 of the Administrative Code without first obtaining a permit from the
of identirpfying and designating species of plants native to Ohioen which are in dangered. Chief of the division. Nothing in this rule shall prohibit the taking or possession of
(Aof extirpation or which are ative Ohio plant specomi es may be designated. species listed on the "United States List of Endangered and Threatened Wildlife
(A Endangered Specif, based on its known statusive Ohio plant species may be designated and Plants" for botanical, education, or scientific purposes, or for propagation in
endafollowing ered if, based on its known status in Ohio, one or more of the captivity to preserve the species, under a permit or license from the United States
following criteria applies:
(1) The species is a federal endangered species extant in Ohio. or any instrumentality thereof.
(2) The natural populations of the species in Ohio are limited to three or
fewer sites, each of which is one square mile or less in area and is 1501:18-2-03. Collecting Permits.
separated from each of the other sites by at least two linear miles. (A) All persons requesting a permit for the taking, transporting or possession of
any native species of wild plants listed as endangered or threatened with
(3) The distribution of the natural populations of the species in Ohio is any native species of wild plants listed as endangered or threatened with
limi3) Th e distribution of the natural populneated by ons of the species in Ohior two United States statewide extirpation for botanical, educational, or scientific purposes, or for
limited to a geographic area delineated by one or two United States propagation in captivity to preserve the species, shall at the time of
Geological Survey 7.5 minute quadrangle maps. propagation in captivity to preserve the species, shall at the time of
Geological Survey 7.5 minute quadrangle maps. application for such permit provide the following information in writing to
(4) The total number of plants in all the natural populations of the species application for such permit provide the following information n wrting to
in Ohio is limited to one hundred or fewer individual, physically (1) A program or project description with a clear statement of study
unconnected plants. objectives.
(B) Threatened Species. A native Ohio plant species may be designated (2) Justificati on of the study.
threatened if, based on its known status in Ohio, one or more of the () Project lcation.
following criteria applies: () Proect location.
(1) The species is a federal threatened species extant in Ohio but not on the (4) Duration of study.
~~~~~~state endangered species list. ~(5) Species and number of specimens to be collected or involved in the
state endangered species list.
(2) The natural populations of the species in Ohio are limited to no less study.
than four nor more than ten sites, each of which is one square mile or (6) Assurance that a final or annual report will be filed with the division,
less in area and is separated from each of the other sites by at least two containing a clear statement of the final disposition of each individual
less in ar ea and is separated from each of the other siteleast two specimen along with a copy of any published papers resulting from the
linear miles.
(3) The distribution of the natural populations of the species in Ohio is study of these plants.
limited to a geographic area delineated by no less than three nor more (B) A permit shall be valid for a period not to exceed one year, but may be
than five United States Geological Survey 7.5 minute quadrangle maps. renewed at the discretion of the Chief.
(C) Rules 1501:18-1-03 and 1501:18-1-04 of the Administrative Code shall be (C) Persons failing to provide full information required in paragraph (A) of this
amended at least biennially to add or delete species to or from the rule to the satisfaction of the Chief shall be denied a permit.
endangered or threatened species lists as new data warrant. (D) Failure to meet the requirements of the permit at the conclusion of the
project shall be reason for the Chief not to issue future permits to the
1501:18-1-03. Endangered Species. The following species of plants are person failing to comply.
(E) A permit must be displayed upon demand to any law enforcement officer
designated in danger of extirpation from the state: having jurisdiction.
having jurisdiction.
(see species list)

APPENDIX F

DATA SERVICES BROCHURE AND APPLICATION FORM

need to acquire written permission to If no information is available on a search
publish information provided by the request, there will be no charge to the data
Heritage Program, and a requirement that user. Otherwise, minimum charge will be
the Division be given credit as the for one hour's work, except where a
information source. memorandum of agreement is involved. Division of Natural Areas & Preserves
The Heritage staff will review the Data Heritage staff will determine the most cost-
Request Form and will contact the efficient search procedures for individual ^1A
prospective data user to answer any data user requests.
questions. A decision will be made about the
level of detail necessary to adequately -iMlll Division of Natural Areas e Preserves
the data user's needs. A cost estimate for
the data search can be given at this time if
requested. This program does not charge for A greemn
data services which result in no data being Division of Natural Areas & Preserves
made available to the user. Charges for Written agreements can be negotiated to
services rendered are made in order to offset fees for services. Data users providing PMP1
recover actual costs to the Division of useftil information to the Heritage Program
Natural Areas and Preserves. can obtain reduced or deferred fees. Contact Division of Natural Areas & Preserves
The requested information will be the Heritage Administrator for details.
retrieved from the Heritage data systems
using the most cost-efficient methods, and
will be provided to the user with specifiede 3 W a
restrictions. If the application for access to Division ofN & Preserves
data is denied, a written explanation will be No prepayment is necessary for
provided, governmental agencies. Prepayment may be
Information will not be provided by required for non-government data users.
telephone or to drop-in visitors. Normally, Invoices normally will be provided along Dierves
data can be provided to users within two with the data. Data users should indicate
weeks of receipt of the Data Request Form. cost limitations they wish to place on
requests for data. Fee estimates can be
given prior to initiating searches.
Division of Natural Areas & Preserves

FeP~~e struaftwe forFOR FURTHER INFORlMATION, CONTACT:
Natural Heritage Program
Division of Natural Areas & Preserves Division of Natural Areas & Preserves
Charges for services rendered are made in Fountain Square Building F
order to recover actual costs to the Division Columbus, Ohio 43224
of Natural Areas and Preserves.
Phone: 614/4668970
Division of Natural Areas & Preserves
DATA MANUAL COMPUTER
USER SEARCHES SEARCHES
Governmental No charge for first $25 base fee +
Agencies - local, hour per month; 2C/line for first
regional, state, thereafter $10 per 300 lines; there- Division of Natural Areas & Preserves
federal. hour. after 1� per line.
Non-Gov't. Agen- $12 per hour. $30 base fee +
cies, Organiza- 2C/line first 300
tions, Individuals. lines; thereafter 1i
per line. JAMES A RHODES ROBERT W. TEATER
Governor Director ODNR
Ohio Department of Natural Resources

Data Services for attractive, but endangered wildflowers should be preserved for scientific and
may lead collectors to these sites. Such educational purposes.
sensitive information is best restricted from REGISTRY - Information on State Nature
In the summernt of 1Natural, the Ohio publication. Preserves; public or privately owned
coDepartment ofTe Natural Resources, in The program's data management system natural areas; and state wild, scenic and
conjunction with The Nature Conservancy, includes manual and computer files which recreational rivers.
established the Natural Heritage Program store pertinent information on rare and OTHER - A catchall class which includes
to systematically identir and help protect endangered species, plant communities and information on bird roosts and rookeries,
significant natural areas, high quality plant unique natural features. This is not a complete breeding animal populations and
glcommunities, rature species and uniquecy listing of all rare species, communities and champion sized trees. These are natural
geologic features. The Naturae Conserancy natural features found in Ohio, but is Ohio's features of statewide significance, but lack
operated the Natural Heritage Program most complete single source of existing natu- legal status.
until October 1977, when the Ohio ral heritage information.
Department of Natural Resources, Division Sound environmental decisions cannot be In what format are the data available?
of Natural Areas and Preserves, assumed made in ignorance of the facts. The Natural The manual, map and computer data
A variety of Heritage Program can provide some basic fms are lexible and allow for searches
information needed to make better use of our syste
organizations contributed biological data to natural resources, while maintaining the which can include a number of variables.
the program, and while this information heritageComputer printouts can be obtained for the
was provided free of charge, a substantial gerations to enoy. following search criteria:
generations to enjoy.
amount of time and money was necessary 1. Geographic Area - County,
to process it into its current useable format. physiographic region, watershed,
In all cases, information was supplied in Wlat data are available? subwatershed, topographic quad, or
the belief it would be used to help protect Heritage data are grouped by class. managed area. Manual searches can
the rare and endangered plants, animals According to the needs of the data user, provide information for areas smaller
and associated natural features of Ohio. In locational information on U.S.G.S. than U.S.G.S. topographic maps or
some instances, locational information topographic maps can be withdrawn for all areas which include portions of
about rare and endangered species was classes, individual class, or parts of classeps.
provided, with the understanding that it The following breakdown summarizes each 3. Species - Individual species, groups of
would not be published or released for class species, taxonomically related species.
public distribution. These agreements of
confidentiality with the individuals and SPECIAL PLANTS (560+ SPECIES) 3. Plant community type - wetlands.
institutions that contributed information * Species with Federal status. 4. Status - Degree of endangerment or
must be honored. * Ohio Endangered and Threatened Plants other special status. (For example,
If data users recognize that the purpose (state official status). legal status only).
of the Natural Heritage Program is to * Plants having no legal status but 5. Date of Information.
protect these species and features, they will considered potentially threatened in 6. Ownership.
better understand why the program staff Ohio.
must carefully control the use of Heritage SPECIAL ANIMALS ( 175 SPECIES) Application Procedure
information. This does not mean the * Species with Federal status. Prospective data users should obtain Data
information is unavailable. In most cases, * Ohio Endangered Wild Animals (state Request Forms from:
data can be provided for specific research official status). Heritage Data Services
projects, development or review of � Animals having no legal status, but Division of Natural Areas & Preserves
environmental impact statements, and considered rare in Oho.
other legitimate uses. However, data users Fountain Square, Building F-
must also realize that publication of exact PLANT COMMUNITIES (65� Community Columbus, Ohio 43224
locational information may not aid the types) Native plant communities of high These forms request detailed information
protection of an endangered species, but quality. about the planned use of the data and
may, in fact, contribute to its destruction. GEOLOGIC FEATURES (40+ Features) Unique require the data user to agree to certain
For example, publication of exact locations or high quality geologic features which restrictions. These restrictions include the

NATURAL HERITAGE PROGRAM
DATA REQUEST FORM

Please enclose: a) A letter formally requesting information from the
Natural Heritage Program.
b) Preliminary reports and maps detailing study area.

DATE

WHO IS REQUESTING THE INFORMATION?

Agency

Name and Title

Address

Phone

WHAT PROJECT OR PROGRAM IS THE INFORMATION FOR? IF THERE IS A CONTRACTING AGENCY,
PLEASE GIVE NAME, ADDRESS, PHONE NUMBER OF CONTACT PERSON.

For which groups in your study area do you want locational information?
Please check appropriate boxes.

Plants -_ Federal Status only
__State Legal Status only (endangered & threatened)
__Potentially Threatened (non-legal status)

Animals -_ Federal Status only
__State Legal Status (endangered) only
__Rare (non-legal)

Plant Communities - _ All __Wetlands only
__Other (please specify)

Geologic Features - All

Other -_ (please specify)

Registry -_ All
__State Nature Preserves
__Natural Areas
__State Wild, Scenic and Recreational Rivers
__Other (please specify)

(over)

-2-

BESIDES NAME, LOCATION, AND STATUS, PLEASE SPECIFY ANY ADDITIONAL INFORMATION
YOU MAY REQUIRE.

DESCRIBE THE LEVEL OF LOCATIONAL ACCURACY NECESSARY FOR YOUR PROJECT.

HOW WILL THE INFORMATION BE USED?

IN WHAT FORM AND DETAIL DO YOU WISH TO PUBLISH THIS INFORMATION?

The information supplied above is complete and accurate. Any material supplied
by the Natural Heritage Program will not be published without prior written per-
mission and without crediting the Division of Natural Areas and Preserves as the
source of the material.

Signature

Return completed form to- Heritage Data Services, Division of Natural Areas
& Preserves, Ohio Department of Natural Resources, Fountain Square, Building F,
Columbus, Ohio 43224. Phone 614/466-8970.