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ANADROMOUS FISHERIES SURVEY OF THE NEW AND WHITE OAK RIVER SYSTEMS 7 i By V) Terry M. Sholar 10 N. C. Division of Marine Fisheries Morehead City, N. C. 28557 ::2: Completion Report for Project A-FC 9 14 November 1975 A QL 628 .N8 S56 1975 TABLE OF CONTENTS INTRODUCTION ....................................................... ..... . I STUDY A.REA ................................................................ 3 MATERIALS AND METHODS .............. ......................... ...... 6 Adult Sampling ........ ....... ...... ......................... 6 Egg and Larval Sampling ........................... ... ............... 8 Juvenile Fish Sampling .............. ................ 11 RESULTS AND DISCUSSION ........... .............. ............... 11 Adult Distribution .... ............................. ............... 11 New River ...... ........... ...... ............. .............. Blueback Herring ... .......... ................... 13' Alewife .............. .................................... 13 American Shad... .... ...... ...... .................... - 13 White Oak River. ...... .......... ................. ........ . 13 Blueback Herring ....... ......................... ..... - 15 Alewife. .......... .... ........ ............ ...... . 15 American Shad ................ ..... ........... . ...... 15 .Other Anadromous Fishes ..... ....... ........ ....... 20 Adult Utilization ......................... ....... ........ 20 New River ....................................................... 20 White Oak River*... ................................... ...... 20 Age and Spawning Frequency... ... .. .. .... ........ ...... ...... 20 New River...... ........................ ....... .... 20 Blueback Herring .................. ........ ..... 20 20 American Shad ..... 22 White Oak River'.... ..... 22 Blueback herring 22 ....................... ..... 22 Alewife 27 Striped Bass ....... ..... 27 Spawning Area Survey ................................................. 27 NewRiver ....................................................... 27 White Oak River ................................................. 31 Juvenile Fish Sampling ............................................... 35 New River ....................................................... 35 Blueback Herring ........................................... 35 Alewife ..................................................... 35 American Shad .............................................. 35 White Oak River .................................................. .35 Blueback Herring ........................................... 35 Alewife ..................................................... 42 American Shad ...................0 ........................... 42 SM91ARY AND CONCLUSIONS ................................................... 42 New River ..................................................... ........ .42 ACKNOWLEDGMENTS .................................I ........................... 47 LITERATURE CITED ...... .......................... ...................... 48 ME .7, V@ fit"! V W"M Z M:@2-, z ?,@'*`5 INTRODUCTION The importance of anadromous; species to the total co rcial finfish landings in North Carolina is shown by statistics of the National Marine Fisheries Service (Table 1). For the 13-year period 1960 to 1972, anadromous species accounted for an average of 46 percent of the total edible finfish landings. A total of seven anadrom'ous species enter North Carolin a waters including the striped bass (Morone ea=tiZie), American shad (AZosa sapidissim), hickory shad (A. mediocris), blueback herring (A. aestivaZis), alewife (A. pseudoharengus), Atlantic sturgeon (Acipenser oxyrhynchus), and shortnose sturgeon (A. brevirostrwn). All seven species are considered both sport and commercial fish and are taken by standard commercial fishing gear. Significant recreational fisheries employ such devices as bow nets, staff nets, dip nets, gill nets, and seines as well as hook and line fishing. The activity provides both products for public consumption and significant recreational and social events affecting large numbers-of resource users. Several studies of anadromous fish stocks "have been completed in North,Caro- h 1-in-A S t-evens dn@`&;--Cl 89 9)' earlier surveyp-zwere- 4p a-in-,a report@-of the: a ad ------- 'Iii- t-1367). sp aua in the Neuse River (Walburg, 1957) and several reports have been-prepared on the Cape Fear River (Davis and Cheek, 1967; Nichols and Louder, 1970). The North Carolina Division of Marine Fisheries began work in the'Albemarle Sound region in 1971 and in the Tar-Pamlico River system in 1974. Baker (1968) published a reconnaissance survey of anadromous fish stocks in the inland waters of North Carolina. Little information is available on the two remaining medium- sized strea- of the southern coastal area: New River and White Oak River. The New River was brieflymentioned in Stevenson's (1899) survey as supporting a shad population. Walburg and Nichols (1967) made reference to the capture of some American shad in the White Oak River. Both the New River and the White Oak River are shown as supporting runs of striped bass, American shad, and river herring in Baker's (1968) reconnaissance of inland fishing waters. Other than these brief general comments, there is virtually no information on the populations of anadromous fishes inhabiting these river system Management of the anadromous fish resources in these river syst requires an information base which can be obtained only through detailed field investi@:,@ gations. The investigations of this project were necessary to provide informa- tion needed to maintain and manage the anddromous fish stocks. Information on basic populations, spawning areas, nursery areas, and utilization ofthe resourc e will - become, vital if the resource base becomes threatened. Objectives of this project were as follows: 1) Job I - To determine the distribution of migrating and spawning adult anadromous fishes. 2) Job II - To determine spawning areas and periods of major spawning activity for anadromous; species. 3) Job III - To determine nursery areas and growth and movements of juvenile -qnndromous fishes. The project was conducted from 1 October 1973 to 30 June 1975. -2- Table 1 Relative importance of anadromous fish in North Carolina as shown by comm rcial landings (from published data: NMFS, Branch of Statistics, Beaufort, N. C.;'N. C. Division of Marine Fisheries, Morehead City, N. C. Total edible Anadromous Percent ana- Year finfish Ubs) fish (lbs) dromous fish 1960 30,470,000 14,308,000 47 1961 30,029,000' 13,490,000 45 1962 31,887,000 16,037,000 50 1963 32,348,000 16,864,000 52 1964 24,562,000 9,183,060 37 1965 33,538,000 14,658,000 44 1966 32,567,000 14,130,000 43 1 1) A 7 @n nRn nnn .71 7-r%n nnr 1968 36,102,000 18,467,0.00 51 1969 41,099,000 22,282,000 54 1970 29,832,000 14,974,000 50 1971 31,380,000 14,991,000 48 1972 48,239,000 13,190,000 27 4@i A -3- STME AREA The New and White Oak River system are located in the south-central coastal area of North Carolina (Figure 1). Both watersheds lie entirely within the lower coastal plain and are mostly swamp drainages with agriculture, forestry, and commercial fishing as the predominant industries. Both rivers are relatively small and drain directly into the Atlantic Ocean in the Onslow Bay area. The New River watershed encompasses about 113,000 ha, lying almost entirely in Onslow County. It drains Wolf Swamp and Hell-and Purgatory Pocosins, then flows southeast for 86 km to empty into the Atlantic Ocean through New River Inlet (Figure 2). New River is the most developed of the two systems with the city of Jacksonville located on its banks. The lower half lies within the Camp Lejune Marine Base. The major tributaries are Southwest Creek, Northeast Creek, Halfmoon Creek, Jenkin-swamp, and Mill Swamp and consist of swamp drain- ages with highly stained waters. The river has an average width of 2 1/2 km., below Jacksonvillesand the average water depth is 1.9--to,2.4 m. The bottom is primarily organic silt excent for sandv shcallareas around the shorelines. Between Jackgonville P-d river kilometer 43.4, the river flows through a hardwood swamp with dense vegetation along its banks where the channel is clogged with stumps and logs.. From river kilometer 43.4 to N.C. Highway 24, a total of 30.8 km, the river was channelized for flood control in 1970 (Figure 2). The lower'13 km of the channelized section is about 15 m. wide, with a depth of 0.6 to 1.8 m, and a bottom type of detritus, sand, and limestone rock. From County Road 1314 to be- low U. S. Highway 258 (5 km), the river is 15 m wide with a rock and sand bottom and is 0.3 to 1.5 m deep. From U. S. Highway 258 to N. C. State Highway 24 (12.1 km), the stream is about 9 m wide with a sand bottom and a depth of only 8 to 15 cm during low flow periods. At N. C. Highway 24, a cement culvert under the roadbed may form a barrier to upstream migration except under high water stages. Some tributaries in the channelized section are connected to the river only through culverts which are not functional for water exchange except-under extreme flood conditions. The river is seperated from its natural flood plain, by dikes approaching 3 m in height. Above N. C. Highway 24, the river is very small and shallow with a sand and mud bottom. The New River system is mostly tidal except for the extreme upper reaches. of the river and its tributaries. Normal stream flow velocities'are very slow. except during flood stages. The White Oak River watershed is slightly smaller than the New River.water- shed and forms the border between'Onslow, Carteret, and Jones Counties (Fignp*.2). It arises in Hoffman Forest and flows approximately 78 km'to empty into the".-,'- Atlantic Ocean through Bogue Inlet. Approximately 77-1,pericent of the wate'rsh6d.- is covered by Hoffman Forest and Croatan National Forest.. There is very.-litile. development in the watershedS with Swansboro being the largest town. The river* is highly stained from swamp drainage. Its major tributaries are Hunter Creek, Grant Creek Pettiford Creek Starkey-Creek,-Black Swamp, and Holston Creek, -4- 78 77 7S COASTAL NORTH CAROLINA 3S- -34 Mot -34 '7H -hai c@P Channe1iied Pottiop of New River. A, 1316 R Quairy. Mck Cr .41116 C, treat er La**& 41 Hunter Cr NO Cr Wallace Cr 010 S gtwe. Iftp. [email protected]*@-.Kerw'@ and White', Oak. Rivir`.- 10 7f, -6- which are small and densely vegetated. The river itself is about 1.5 km wide from the Atlantic Ocean to the vicinity of Webb Creek. The depth is about 1.2 m. with numerous oyster reefs and a narrow obscure channel. The bottom type is primarily sand and mud with sand shoals along the edges. From Webb Creek to Crant Creek, the river is about 3.7 m. deep, 150 m. wide, and meanders. The bottom type is primarily mud and detritus and is boardered by fresh and brackish marsh. Above Grant Creek, the river flows through hardwood swamp at a depth of 1.5 to 4.3 m. Below U. S. Highway 17, the river flows through seven distinct lakes of the Martin Marietta Belgrade Quarry. They comprise approximately 60 ha of water and range from 4.5 to 9.5 m in depth with steep sides and mud bottom., The lakes were dug between 1940 and 1960 during mining operations for limestone. Water flow is very slow in the lakes except where they are connected. From the Quarry to immediately above U. S. Highway 17, the river is narrow'and from 0.6 to 1.2 m deep with a rocky bottom. The flow is very swift in riffles areas. Above U. S. Highway 17, the river is a small typical swampy stream. The White Oak River system is predominantly tidal except for the region above the limestone quarry and the headwaters of the tributaries. The stream. velocity is slow, but is greater than in the New River watershed. A detailed survey and description of the New and White Oak River systems ' was prepared by the North Carolina Wildlife Resources Commission (Davis and McCoy, 1965). MATERIALS AND METHODS lutial sampling indicated three basic ecological zones*in each river consisting of high salinity, brackish, and fresh areas (F 'igure 3). The high salinity zone consisted of the lower portion of the estuary and generally coincided with the open water areas. The brackish zone consisted'of a transition area between the lower estuary and the fresh water zone. The fresh zone cou- tained swamp areas exhibiting little or no salinity. Sampling and data recording was done on the basis of the three zones. The time between 1 October 1973 and 30 June'1974 was'an initial survey period. Sampling during this time was generally random witIt no permanent stations. Permanent stations were establishe4 and sampled regularly between I July-1974 and 30 June 1975. Advit:S -lin Adult anadromous fish sampling was done on the-basii'of high'deais; ty-,-.-ra'ndo'x' sampling b etweein- I January and 30- June 19 74 and, ripire4imtsUme. -stitious IIIVere@, sampled betweem",'I Januaiy and 30 june.1975 with kiU. -..net4 "and '.ii 'nets @I 7 5,03. AP -7- High salinity zone-- Brackish zone --- - ---- Fresh zone ------- 7--- 0 N3%A, 1316 rs 17 A Quarry. Cr S 'Cr CP Cr at e Cr A? F reemaltr Hunter Cc Wallace Cr 0-0 .00 --acque Inlet Iniet. R"Ve 7 zones in-- ihi': New and -White.` Oak Rivers-..--. Figure 3 Map,:of the 'thrde@ -8- (Figure 4). Gill nets of 45.9 and 9.2 m legths with mesh sizes of 6.4 cm, 7.0 cm, 8.3 cm, 10.2 cm, 12.4 cm, 13.3 cm, and 14.0 cm stretch mesh were used to sample adults. The 45.9 m nets were used in the high salinity and brackish zones, while the 9.2 m lengths were used in the fresh zone and the tributaries. There was no high salinity zone sampling after March 1975. *Trammel nets of 91.4 m lengths were used primarily to sample striped bass-and sturgeon. All nets were anchored and left for a 24 hr period. Catch-per-unit-of-effort (C/E) was calculated on the basis of a Standard Fishing Unit (SFU) of 91.4 m. of net set for 24 hours. The C/E for herring was based on the number of SFU of small mesh sizes, and shad C/E was based on the units of mesh sizes 10.2 cm and greater. One trammel net effort was based on 91.4 m of trammel net set for 24 hours. The adult fish were sorted by species, weighed in grams, and fork lengths (FL) were measured in millimeters. The gonads of each individual 'were examined to determine sex and stage of development which was recorded-as immature, mature, ripe, running-ripe or spent. Scales were taken from each individual and placed in envelopes and coded along with sex, length, and weight information. A lar e number of scales were taken from below the front of the dorsal fin. Scales were read with a binocular scale edge counted as a yearmark, as described by Marcy (1969), Beal (1968), Cating (1953), and Street and Adams (1969). Several clear scales from each fish were examined. Two readings of each set of scales was made, and those not agreeing were discarded. Egg and-Larval Sampling Sampling for eggs and larvae was conducted between 1 March and 30 June 1974, and 17 March and 30 June 1975 with I m and 1/2 m plankton nets of 00 mesh. Where possible, a station was sampled for each five mile segment of river and major tributary (Figure 5). Sampling was conducted twice monthly in each river alternating weeks between rivers. All samples were taken within 10 cm of the bottom and/or within 10 cm of" the surface. Towed samples were of 5 -minute duration and sets from bridges: were of 10 - to 15 -minute duration depending upon the current velocities NO eggs, were captured in any 15 minute sets so catch-per-effort was based on a 5-minute tow. Results of the I m and 1/2 m nets were treated separetely. "Eggs and- larvae Samples were preserved in the'field with 10% of formalin. Eggs and larvae were sorted,identified, and counted in the laboratory.- alewife eggs and early larvae were lumped together as "river" hearing due to an Inability to distinguish between the two species -9- Adult sampling stations 0 Roo -1316- 17 Oudr* ry dO X Cr 1116 11 U.1 ell Ist Lake Wallace Cr 0 010 Pet ivet 5,09us lnl*t Inle I A-. @iivers :-'1975 Fi' .4 Iftp.- -of adult. siavOlilig--s-tatious New.-wid White 0. gure Egg sampling stations 13N Ran 31 17 All. r Cr F ret man CO. Cr pel t- 1369us infol OT 1jtjV e Vi 14. Oak.- Ri*ei6 _,Stat and te, Whi irk --vigitr @99: zXy'@t zts;e L, Z71. -A R d. '1-1,5511@ 3- Juvenile Fish Sampling Juvenile fish sampling was conducted from 1 October to 31 December, 1973 and from I July to 31 December,1974. Initially, modified Cobb pelagic trawls of 1.8 m. and 3.7 m width were used in open water areas and shoreline areas not suitable for conventional seining. Results showed the Cobb trawl to be selective and inefficient for capturing juvenile%anadromous fishes. A modified bottom trawl also produced unsatisfactory results. A modified wing trawl of 1.9 m head rope length was designed and tested in July of 1974. This gear showed the most satisfactory results and was subsequently employed as the standard gear. It had a mesh size ranging from 10.2 cm in the wings to 0.63 cm in the tail bag. All samples with the wing trawl were taken on the surface by using surface doors. A standard effort with the modified wing trawl consisted of one 10-minute tow All other samples were taken with 0.63 cm mesh bag seines of 18.4 m. and 6.1 m lengths. One haul with a 18.4 m or 6.1 m seine constituted one 18-4 m or 6.1 m seine effort. All samples were presetvid in@the field with 10% formalin and were later 6ol@p_@as - to--species, weighed t o the nearest.. Igr4m,.and fork lengths measured in-- LUALIILL@@ @6. A tiL;UtiaLU1j1e 01- ju xisix was Laken itL_!a-k0:e s amp les. _-Incidentai species such as spot, croakers, sunfish, etc. were counted, weighed,.measured, and recorded as to species. Random sampling was conducted between 1 October and 31 December, 1973 primarily using the 18.4 m seine. From 1 July to 31 December, 1974, sampling was done regularly at established stations (Figure 6). Each river was sampled two weeks per month alternating weeks between rivers. There were a total of 26 stations in the New River making 52 samples per mouth. The White Oak River had 25 stations resulting in 50 samples per month. RESULTS AND DISCUSSION Adult Distribution New River A distribution of adult anadromous fishes was determined by employing a total of 219.75 SFU of gill net effort in both river systems. A total of 480 adult anadromous fishes was captured. Blueback herring and alewife were Mo 'st numerous with fewer numbers of American shad, hickory shad, Atlantic sturgeon, and striped bass. During 1974 and 1975 in New River, a total of 95.75 SFU of gill net effort, and 9 tramnel net sets produced a total of only 66 anadromou's fish, for an average catch of 0.7 fish per gill net SFU. No adult anadromous fish were caught in any trammel net samples. The catch consisted of 46'blueback herring, 7 alewife,'9 American shad, and 4,Atlantic sturgeon. 5 -12- ------ ---- Wing,Trawl- 6.1 m Seine-1- ---------- 18.4 m SE@_ine ------ - ----- 0 qzl 13-6 Roo mo Quor ey alaik Cr 'AMS great take freemv, Hunter Cf Cr 1Wallace Cr is* Fri, 70 0 9 04P Inlet-, tei d Ri of 17elln f-iih sac4 _g@sta ;2- @rg 5 R 1 ZZ, V 13 Blueback herring Eight adult blueback herring were captured in the New River during the run of 1974 (C/E=0.3). They first entered the river in January with the peak occurrence in the fresh zone in April. In 1974, they were found up to the lower channelized section of the river where they were observed to be in a ripe condition. None were taken in the tributaries, and by the end of April no adult blueback herring were seen in the river. In 1975, 38 blueback herring were captured (C/E=22). They were found in the swamp below the channelized part and in certain tributaries. Only one was caught in the channelized section. Blueback herring were observed, using North- east Creek, Little Northeast Creek, Blue Creek, and Hawkin Creek. Hawkin Creek had the highest single C/E which was 5.8. Blue Creek and Little Northeast Creek had a higher C/E than the main run of the river (Table 2). One adult male alewife was caught during the 1974 run in New River. During the 1975 run five alewife were found in Blue and Hawkin Creeks. Blue Creek contained a larger run with 2.4 catch per SFU (Table 2). Their peak occurrence was during January and February. American shad In 1974, four adult American shad were captured for a C/E of 0.3. They were taken up to County Road 1316 and were in a ripe condition. None were found. in the tributaries. During the 1975 run, four American shad were captured for a C/E of 0.3. None were found in the fresh zone of the river where they occurred in greatest numbers the previous year. Three of the four were found in Southwest and Little Northeast Creeks (Table 2). They were found in the river system from February through April. No adult striped bass or hickory shad were captured in the New River. A single Atlantic sturgeon was captured during March, 1974 while three were caught during March and April of 1975. It is apparent the New River system has limited anadromous fish resources. Only a few blueback herring, alewife, American shad, and Atlantic sturgeon were found in the river. A relocation of the distribution of anadromous resources appears to be occurring. They appear to be switching, from the upstream, utilized in 1974, to the downstream tributaries used in 1975. This could be due to alter- ations caused by channelization of the upstream area. White Oak River In the White Oak River, a total of 124.0 SFU of gill net and seven trammel net sets produced a total of 636 anadromous fish for an overall gill net catch ` of 5.1 fish per SFU. No adult anadromous fish were caught in any trammel net -14- Table 2 - Catch per Standard Fishing Unit for adult blueba'ck herring, alewife, and American shad by area% New RiverP 1974-75, Catch per SFU Species Area .1974 1975 Blueback herring New River 0.3 1.1 Northeast Creek 0.0 1.0 Little Northeast Cr6ek 0.0 2.5 Blue Creek 0.0 1.2 Hawkin Creek 7.9 Alewife New River 0.1 0.0 Blue Creek 0.0 2.4 Hawkin Creek -0.0 0.8 American shad New River 0.2'@ 0.0 Southwest Creek 0.0 0.1 Little Northeast.Creek:... 0.0 1.3 t g -15- samples. The catch consisted of 487 blueback her-king, 90 alewife, 49 American shad, 2 hickory shad, 3 striped bass, and 5 Atlantic sturgeon. The average catch per SFU for blueback herring was 7.3; for alewife, 1.4; for American'shad, 0.9; for hickory shad, 0.04; for'striped bass, 0.03; for Atlantic sturgeon, 0.04. Blueback herring In 1974, 177 adult blueback herring were captured for a C/E of 5.3. They were found in the White Oak River up to the Quarry Lakes and occurred in the river from January through May. None were found in the tributaries. During the 1975 run, 310 adult blueback herring were captured for a C/E- of 10.3. This suggests that the run of 1975 was about twice cis large as the 1974 run. Again they were found up to the*Quarry Lakes and occurred in the river from January through May (Figure 7). Bluebadk he-ki-ing were also found in Hunter, Webb, Grant, and Holston Creeks. Unlike the New River the catches were small compared to the main stem of the river'. Holston Creek had the highest number of blueback herring in any tributary with a C/Eof 2.2 compared to an 11.4 C/E in the main stem of the river (Table 3). Alewife In 1974, 10 adult alewife were caught in the White Oak River for a C/E.of 0.3. They occurred during April and May up to the'Quarry Lakes'. None were found in the tributaries. Adult alewife did not*enter'the river until March or April, well after the first blueback arrived. This was different from the reports of alewife arriving earlier in the Albemarle Sound area by Street and Pate (1975). During the 1975 run, 80 adult alewife were captured for a C/Eof 2.4, indicating a much larger run in 1975. They were found from February through May:. up to the Quarry Lakes (Figure 8). Alewife were also ciaught'in Hunter, Grcmt*,. and Holston Creeks. They occurred'in slightly greater numbeiik in the tributaries than in the river*. Grant Creek had the'highest number'with a C/E.of 3.7 to a C/E of 2.8 in the main stem of the river (Table@3). American shad Twenty-one adult American shad were caught in 1974@ for a C/E. of 1.2,' They occurred from February through May up to- the Quarry Lakes. None were found in the tributaries, In 19 75, 28 American shad were found for' a C/E. of 0. 4 indicating a - smal,lef run than in 1974. They again occurred' from'February through May up tbAhe-Quarry Lakes (Figure 9). Pettiford Creek was the'6nly tributary containing American'. shad with a catch of 0.7 per SFU. I'M -16- 50" High Salinity Zone 40- 1974 30- - - - 1975 c /e 20- 10- 0- Jan Feb Mar Apr May June Brack'ish Zone 197A 40- 30- c /e, 20- 10- 0, Jan Feb Mar Apr May June So- F r e s h-Zone 40- "30 20 nf '10 j 7r S@ 27 F 9 Im -,:May :@_@pUwnp' @R, 4974-75 -17- Table 3 - Catch per Standard Fishing Unit of adult blueback herring, alewife, and American shad by area, White Oak-,-River 1974-75 Catch per SFU Species Area 1974 1975 Blueback herring White Oak River 7.3 11.4 Hunter .Creek 0.0 1.3 Holston'Creek 0.0 2.2 Grant Creek 0._0 0.7 Webb.Creek .... .. ....... 0.0 1.6 Alewife White Oak River 0.3 2.8 Hunter*Creek 0.0 0.6 Holston Creek 0.0 3.4 @n'_n American shad White Oak River 1.7 0.8 Pe ttiford Creek 0.0 0.7 2 0 firackish Zone -1974 .---1975 1 5 c/e 5 0 Jan Feb Mar Apr May June 20 Fresh Zone -1974 - - - 1975 C A 10 5 0 Jan Feb Mar Apr :-,,:May It Z4@ gure k53 451 t'. 7"z: -@7- R. ffil -k. _19- 7 _Orackish Zone -1974 .6 - - - 1975 5 4 c 4 3 2 0 Jon Fe b Mar Apr May June Fresh Zone 7 -1974 6 - - - 1975 5 4 c Ae 3 2 0 -4on .-Feb Mar Ap r.@. ...MaY 19 74-75 -per- SM-- of -adult _Americim: sbuj&-by --year, _.@_white-Oak. :River'*'.. - Figure __77 --L4 L -20- Other anadromous fishes Two adult hickory shad were found in the White Oak River. One was caught in January 1974 and one in May 1975. While no striped bass were found in 1974, three were captured in March of 1975 in the lower part of the river. Five Atlantic sturgeon were found in April of 1975 in the lower part of the river. None were captured in 1974. Adult Utilization New River The New River contains little or no anadromous fishery. Only a few American shad and herring were caught incidental to other fishing operations and were usually sold. The value is extremely minor compared to the overall New River fishery. White Oak River The White Oak River supports a recreational dip net and gill net fishery mainly. for herring. Fishing is primarily confined to the Quarry Lakes and may involve 25-30 local people per night with catches of 5O-100 pounds per person during the peak. The incidental catches of anadromous fish by commercial fish- ermen is minor, with little value to the White Oak River fishery. Age and Spawning Frequency New River Due to the small number of adult anadromous fish sampled in the New River, no definite conclusions can be made as to their age or spawning frequency. Blueback herring Only thirty adult blueback herring had scales suitable for aging. The ages of both males and females ranged from four to six years with. ages four and five dominating. The mean fork lengths for each age group generally agreed with those found by Street and Pate (1975) in the Albemarle Sound area (Table 5). However, due to the small sample size from the New River, no adequate comparison can be made. The ratio of males to females was about equal. Spawning repetition by blueback herring in New River was extremly low. Only 10 percent contained spawning marks which differs greatly from the 50 percent found by Street and Pate (1975). None were found containing more than one spawning mark (Table 6). Alewife Seven alewife from the New River had readable scales. Ages were four and five, with age five predominating. All males were age five while females were -21- Table 5.-Total number, percent of total sample, mean fork length, and length range for each age group by sex for blueback herring, New River, 1974-75 Total Percent of Mean fork Length Sex Age number total sample length(mm) range(mm) Female IV 7 50 246 240-253 V 7 50 249 237-255 14 Male IV 9 56 233 226-246 V 5 31 250 241-256 VI 2 13 268 267-268 16 Table 6 -.-.Age and spawning frequency of blueback herring from New River, 1974-75 No. times spawned 0 1 Total Age 14 F M F IV 9 7 0 0 9 7 V 4 6 5 7 V1 0 @l Or 21. Total 14 13 .2." 14 @Percent- (88)@ @(93) (1-3) (7) 10 li56 -22- ages four and five. The fork lengths were not comparable to those found by Street and Pate (1975), probably due to the small sample size (Table 7). Eighty- five percent of the-alewife were repeat spawners. One virgin female was found while all males had spawned previously. None were found to spawn more than once. American shad Five American shad from the New River proved suitable for aging. The males were age four and five while all of the females were five. Their mean fork lengths were not comparable to those found by Street and Pate (1975) (Table 8). All were found to be virgin. White Oak River blueback herring Of'the blueback herring scale samples taken in the White Oak River, 318 could be read. They ranged in age from three to seven years with 91 percent of the males being ages four and five, and 57 percent of the females being age five. The# fork length agreed closely with those found by Street and Pate (1975) in The male to female sex ratio was 0.88:1, quite different from that found by Street and Pate.(1975). This is probably due to the selectivity of gill nets used in this study. The number of virgin blueback herring was higher than that found in the Albemarle Sound area with 79 percent being virgin. Females were observed to re- peat more than males. Only four females (1.3 percent)-had more than one spawning mark. Males first spawned at ages four and five while most females first spawned at age five (Table 10). Alewife Fourty-two scale samples from alewife were useable. Males ranged in age from three to five years with 72 percent being age four. All females four and five, with age four predominating. The fork lengths agreed With those found by Street and Pate (1975) (Table 11). The number of virgin fish was higher than expected, With 92 percent virgin males and 94 percent virgin females. None were foRmEd to have spawned more than, once. Both males and females were found to first spawn predominately at age- four (Table 10). Of the alewife found in the White Oak River, 60 percent were males and 40 percent females. This Sex rationdiffers greatly from that expected, but the samplesis so small that no, conclusions can-be made. American shad A total of thirty-five scale samples of American shad were suitable for aging. Females ranged in age from four to seven years with age five making up 56 percent. -23- Table 7 --Total number, percent of total sample, mean fork length, and length range for each-age group-by sex for alewife, New River,' 1974-75 Total -Percent of Mean fork Length Sex Age number total sample length(mm) range(mm) Female IV 1 33 240 V 2 67 289 280-297 3 Male V 4 100 266 262-267 Table 8 Total number, percent of total sample, mean fork length, and length @ranges fok eachage.-group, by sex for Americah.'shadj. MO-w-River, 1974-75- Percent of Mean fork Length Sex Age. numb eir total sample length(tam) range(MM)- Feviale V 3 100 465' 441-498 Male IV 1 50 398 V I so 387 -24- Table 9 --Total number, percent of total sample, mean fork length, and length range '@''@_for@each`age group..by sex for blueback herri.ng,..WUte'.Oak,River, 1974-75 Total Percent of Mean fork Length Sex Age number total sample length(mm) range(mm) Female IV 38 23- 242 202 - 262 v 97 57 258 234 - 277 vi 31 18 265 235 - 281 vii 3 2 280 260 - 281 169 Male 111 2 1 219 208-- 229 IV 68 46 237 223 - 253 IA7 11-7 -171) vi 12 8 249 236 - 269 149 X iz @Z R; Zt -25- Table 10 - Age and spawning frequency of blueback herring, alewife, and-American shad from White--Oak River, 1974-75 Blueback herring No. times spawned 0 1 2 Total Age M M, F M F., M F 2 0 0 0 0 0 2 0 IV 63 37 5 1 0 0 68 38 V 51 77 16 19 0 1 67 97 VI 5 14 7 @14 0 3 12 31 VII 0 1 0 2 0 0 0 3 Total 121 129 28 36 0 4 149 169-,' Percent (81) (76).,@,' (19) (21)_ (3)_. Alewife No. times s2awned 0 1 Total Age M F M F M F 111 2 0 0 0 2 0 IV 16 12 2 1 18 13 V 5 4 0 0 5 4 Total 23 16 2 1 25 17 Percent (92) (94) (8) (6) Ameikican -shad No. times spawned 0 1 Total ,Age M M F IV 10 2 0 0 M 2 V 8 9. 1 0 Vi 0 4 0 VII 0 1 0- 0 0 Total 18 16 1 0 19 16 Percent (95Y., (100), -26- Table 11 - Total number, percent of total sample, mean fork length, and length range -.for each age group by sex.for alewife, White Oak River, 1974-75 Total Percent of Mean fork Length Sex Age number total sample length(mm) rauge(mm) D-male IV 13 76 256 249-266 V 4 24 272 260-289 17 Male 111 2 8 231 225-237 IV 18 72 241 225-261 V 5 20 250 242-'255 25 Table 12 Total number, percent of total sample,*mean fork length, and length range - for each age group,by:s6x for-American shad,. White'.Qak Riveri 1974-75 .Total .... Percent of Mean fork Length Sex Age number total sample length(mm) ranze(mm) Female IV 2 13 417 391-442 V 9 56 474 452-485- VI 4 25 481 467-492 Vil 1 6 520 16 53 Male IV 10""'. 406 -363431 V 9 47. 449 '416-482 19- J! 4 q -27- Their fork lengths agreed or were slightly smaller than those found by Street and Pate (1975) (Table 12). All were found to be virgin except one five-year old male which had spawned once previously (Table 10). This is currently the southern-most confirmed report of an American shad repeating. Hickory shad Two male hickory shad were caught in the White Oak River. One 345 mm three-year-old was found in January, 1974, and one 318 mm, four-year-old was found in May, 1975. Both were virgin. Striped bass Three striped bass were found in March, 1975. Two were three-year-old females measuring 394 am and 401 mm. The other was a 712 mm seven-year-old female. Spawning Area Survey New River Spawning in the New River appeared'to be very slight. In 1974, only two river herring eggs were taken in 8 samples using the'l-meter egg net and 100 samples using the 1/2-meter egg net. They occurred in only one sample during the week of 6 May below County Road 1316. This indicated the lower channelized section as being the spawning grounds for river herring (Figure 10). Five hickory shad eggs were captured during the week of 20 May below N.C. Highway 24. Spawning was probably limited to this point by'the barrier'presented by the culvert under Highway 24 (Figure 11). No American shad eggs were captured, but one rimning-ripe-female taken in the channelized area indicated that some spawning occurred in the altered section*of the river'-(Figure-U).@ No anadromous fish larvae were.taken. During 1975, 69 river herring eggs and two larvae were taken in 95,.1/2- meter egg net samples. No river herring eggs were found in the river' * Sixty- six eggs were caught in Hawkin Creek and three were found in Blue Creek-during Nay. Two herring larvae were also found in Blue Creek during May (Figure 10)., No running-ripe female herring were found. Noeggs or larvae of other anadromous fish were found. However, one running-ripe female American shad was taken in Little Northeast Creek indicating it to be .4 spawning area. Although no eggs or runnin&-riperfemales were found in Southwest Creek the*presence of adults indicates American shad possibly spawn there. Also, the presenize 6f'adult.blue- back herring indicates these possibly spawn in Northe"t and Little Northeast:- Creeks. All o@jj-neittiug- results indicate very poor.spawning succe"-for all an-a- dromous species in New River. The'prior'loc'atiou'of'spawni.ng grounds :Es not.- known; however, the section*of'river presently channelized'does''not',appeak to offer suitable spawning habitat. -28- area----- Spaw@iing 7 Probable spawning area-, ,0 lRd "all arry Lake FtemV, wMace Cr 00 ell cr "White vak Riv-ers 1974-75. fare".. ror., lq@- j@p Pf,,- 13pajrn$, rivim'herrinj :J 7;@- -29- Spawning area----- 1316_. 0 a Quarry OS Cr z fp ell cr tail fr!tmaar Cr Cr pellifer Cr Rogue 1111*t Inlet sew R 'Figure 11 Hap of spawning,it a f6 hickory. 4had- New River,1974 re r -30 59 Spax@ain gi area zi Possible spawning area 17 c121 Alit ucrry c". Like, Z freeman r 94 or cc Wallace Cr 010 Vey of spairniqg air" of :Amri@an, A@a -New- ai2d'whitel Oak R:Evikd'@-.z-19,74 .. 75 Map J. _4 -31- White Oak River During 1974, a total of 2,083 river herring eggs were taken in a total of 10 samples using the 1-meter egg net and 105 samples using the 1/2-meter egg net. All were found in the Quarry Lakes. No other anadromDus fish eggs were taken. River herring began spawning the week of 2 April and extended through the week of 13 May. Peak spawning occurred during the week of 29 April (Figure 13). During 1974, a total of 45 river herring larvae were taken in a total of 41 samples using the 1/2-meter egg net in the fresh zone of the White Oak River. They were caught from 2 May to 15 May in the Quarry Lakes. All were early pro- larvae and had a mean length of 3.5 mm. The peak capture of river herring larvae occurred on 15 May (Figure 14). During May, 90 percent of the blueback herring found in the Quarry Lakes were running-ripe, indicating this to be a major spawning area (Figure 10). No other anadromous fish eggs or larvae were caught during 1974. The capture of a running-ripe female American shad in the Quarry Lakes indicates this to be a spawning area (Figure 12). No r-ning-ripe female alewife were -fo-und-, but their presence in the Quarry Lakes probably indicated this to be a During both years, spawning of river herring was observed taking place at the Quarry Lakes below U. S. Highway 17 and in the shallow',, rocky region above the Quarry. Several schools of herring were observed "washing" along the edges, where the swift current produced rapids as it flowed over rocks ' Herring were seen ascending a small waterfall (0.5 m. in height) and spawning was observed taking place above the fall. During 1975, 891 river herring eggs, 8 river herring larvae, 28 blueback herring larvae, 3 alewife larvae, and 1 American shad egg were captured in 87, half-meter egg net samples. River herring spawning began the last week in March and lasted through May with peak spawning occurring during the first week in May (Figure 13). Spawning was observed in the Quarry Lakes as In 1974, but evidence also showed that - spawning occurred in certain tributaries. River herring larvae found in Holston Creek and Grant Creek indicates these to be spawning areas (Table 13). The presence of adult river herring indicates spawning probably also occurs in Webb and Hunter Creeks, although no positive evidence was found. American shad were found to spawn in the Quarry Lakes as indicated in 1974 (Figure-12). One egg was captured immediately above the*Likes during the first week in May. Also, runnin&-ripe female American shad were obs'erved in 1975 in@ the Lakes. The presence of adults indicated that spawning probably occurred'in. Pettiford Creek Vdit's piositivwsevidence vm fotmd. -32- c Ae 275 - 1974 -Tomperatur 250- 225-- -25 200 - 175 20 3 c /e 150- V a 125- 15 c 100- 75 - 10 0 50 25- -5 0 0 4/2 4/16 4/29 5/13 5/28 W k c/e 1975 --Temperature 50- 25 40- -20 30 - 14 c /k 20- .10. :..c to 10- a 5 0 0 @3/24 417 4/21 -5/5 w 6 0 k AR VISW im :-7 9 _e Fi z, -33- 1974 4 3 c /e 2 0 4/16 4/29 5/13 5/28 Week 5 1975 4 3 c /e. 2 0 L 4/7 4/21 5/5 5/19 W o e k --hem Figure 74aVe-E-Z-1 ORM -34- Table 13 - Evidence indicating spawning area locations, White Oak River, 1975 Location Date Species Comments Quarry lakes 3/26/75 River herring 112 eggs captured 3/26/75 Blueback herring Running ripe female-gill net 4/10/75 River herring 20 eggs captured 4/24/75 River herring 24 eggs captured 4/24175 Blueback herring Running ripe female-gill net 4/24/75 American shad Running ripe female-gill net 5/7/75 River herring 672 eggs captured;..spawning activity observed 5/19/75 River herring 63 eggs captured; spawning activity observed 5/19/75 Blueback herring Rimming ripe female-gill net 5/19/75 American shad 1 egg captured Holston Creek 5/5/75 River herring 1 1@._ captured 5/5/75 Blueback herring Running ripe female-gill net 5/20/75 Blueback herring 24 larvae captured Grant Creek 5/6/75 River herring 7 larvae captured 5/20/75 Alewife 1 larva.@, captured -35- Juvenile Fish Sampling New River A total of 90, 18.4 m seine efforts; 122, 6.1 seine efforts; 43, 1.8 m Cobb trawl efforts, and 130 wing trawl efforts produced 9 blueback herring, 2 alewife, and 2 American shad from October through December, 1973 and July through December, 1974. Catch data for each gear type are shown in Table 14. The 18.1 m seine and the modified wing trawl were found to be most successful at catching Juvenile anadromous fish. The 6.1 m seine was used in the smaller tributaries and produced no juveniles. Blueback herring Juvenile blueback herring were found during October and November, 1973. None were captured during 1974. All of the blueback herring were found in the lower channelized section (Figure 15), and had a mean fork length of 51 mm in October 1973. The data obtained was inadequate to determine growth and movement. Alewife Only two juvenille alewife were found in 1973 (Table 14). Both were caught n the lower channelized section during October 1973 (Figure 15). None were American shad Two Juvenile American shad were caught in the New River (Table 14). One was taken in October 1973 in the lower channelized part and one in October 1974 in lower Southwest Creek (Figure 16). This may indicate Southwest Creek as a possible nursery area. Juvenile fish sampling indicates that the New River produced very few young- of-the-year anadromous fish during 1973 and 1974. The small catches of juveniles may be a result of reduced spawning and a lack of suitable nursery area habitat to maintain an anadromous population, but it will be very reduced. White Oak River From October through December, 1973 and July through December, 1974, a total of 352 blueback herring, 110 alewife, and 3 American shad were caught in 159, 18.4 m seine efforts; 81, 6.1 m seine efforts; 30, 1.8 m Cobb trawl efforts; and 83 wing trawl efforts. The C/E by gear is shown in Table 15. Blueback herring Juvenile blueback herring were found from July through November in the fresh zone of the White Oak River. Their peak occurrence was in October' of 'both yen s, with catches droqpping drastically afterwards (Figure 17). This indicates the seaward migration probably occurs after October. Three tributaries produced Juvenile blueback herring. They were Webb, Grant, and Hunter Creeks with Hunter having the highest C/E(Figure 15 and Table 16). -36- Table 14.- Catch, effort, and catch-per-unit-effort (C/E) for 1973 and .1974. year classes of blueback herring, alewife, and;American shad-by' gear,'Ne-w-River 18.4m, seine Wing trawl 1. 8fiL Cobb trawl Year 1973 1974 1973 1974 1973 1974 Effort 36 54 130 43 Blueback herring Catch 9 0 0 0 C/E 0.25 Alewife Catch 2 0 0 0 @C/E 0.06 American shad Catch 1 0 1 0 C/E, 0.03 0.01 -37- Nursery area Roo 13i6-- It, Quarry. We S p ood treat Lake Freema W*ter Q Wallace Cr peltilord 71 809we 'RI'my Figure -15:.'-;- -Map of:nursery area of.blueback.@herking-and aldwife... N.dw and White-Oak River- -38- Nursery area @Loa Roaa 1316 Quar'ry* Cr FIl 0-p Great Lake Hunter Cy Wallace Cr AV 0 Pe 8051 u's Inlet it. Inlet Ak Rivie-ts can -16 NO a@dd @idh"6 0:. Tigure' mip :of -m@uraiery 'area Of Amii dhAd -39- Table 15 - Catch, effort, and catch-per-effort (C/E) for 1973 and 1974 year classes of blueback herring, alewife, and American shad by gear in the White Oak River 18.4m seine Wing trawl 1.8m Cobb trawl Year 1973 1974 1973 1974. 1973 1974 Effort 36 123 83 30 Blueback herring Catch 224 87 35 6. C/E 6.23 0.71 0.43 -0.20 Alewife Catch 1 42 67 0 C/E. 0.03 0.35 0.81 American shad Catch 0 3 0 0 C/E 0.03 -40- 14 1 2 10 8- -18.4-n Seine C /e. 6- 4 2- -No,v---..Dec Jul Z-Aug- Sep. Oct ---Nov Doc -er-f owictue, Figure I-T, :1973-74,-.- -7-t Z7@ _4 -41- Table 16 - Catch-per-unit-of-effort for blueback herring and alewife by gear and area, White Oak River, 1973-74 18.4 m seine Wing trawl Year 1973 1974 1974 Blueback herring Brackish zone 0.7 0.0 0.0 Fresh zone 11.6 1.2 0.3 Hunter Creek 0.9 Webb Creek 2.0 Alewife Brackish zone 1.0 0.4 Fresh zone 0.1 0.4 1.9 Grant Creek 0.1 Hnlqton Creek 0.6 Webb Creek 0.1 Hunter Creek 3.6 The mean fork lengths by month for blueback herring were larger than those found by Street and Pate (1975) in the Albemarle Sound area*(Figure 18). This is probably due to their use of 3.2 mm mesh gear as opposed to the 6.4 m mesh gear used in this study. Alewife Juvenile alewife were found in greatest concentrations in the fresh zone in September. Afterwards, their numbers decreased ' indicating the seaward migration occurs after September (Figure 19). The nursery area for alewife generally corresponded with that for blueback herring (Figure 15), but with a higher concentration of individuals occurring in the lower part of the river (Table 16). They were also found using the lower parts of Webb, Hunter, Grant, and Holston Creeks with Hunter Creek containing the highest concentration (Table 16). The mean monthly fork length for alewife ranged from 47 mm in July to 81 mm in December. This agreed with those found by Street and Pate (1975) in the Albemarle Sound area (Figure 20). Juveni le Ameri can shad were taken only in July of 1974. It appeared they left the spawning grounds and immediately moved down stream on a seaward mi- gration. They were found from the spawning grounds down to the vicinity of Holston Creek during a one week period in July (Figure 16). At this time the mean length of the Juvenile fish was 59 mm. The White Oak River apparently produces enough Juvenile anadromous fish to maintain an exploitable population. The nursery grounds of the three species combined, covers over 30 km of river from the Quarry Lakes to the Webb Creek area. SUMMARY AND CONCLUSIONS New River The New River appears to contain very low anadromous fish-i3tocks. As a result, there is little or no utilization of anadromo'us' fish-. The reduced pop- ulation is probably due to the"30.8 km'of'channelized stream. and the large amount of development of the New River watershed. Personal communication with commercial fishermen indicated significant anadromous populations once existed. The drastic reduction'in anadromous fishes in the'Nev'River is probably due to reduced recruitment. Low spatf4ng and Juvenile. success as seen in 1973 -And 1974 is probably due to a lack of suitable habitat which'i),laces'thi Anidrom6us,- stocks in serious danger'. The'future of th6'Nev River'--adromfoUs' population will depend upon the Ability- of the'tributaries to provide suitable habitat to- maintain recruitment since the habitat previously found in Nev'River'above' Jacksonville no longer exists- -43r 72 - 70 - 68 - 66- 64 - 62 E E 60- -C 58 56 54 b. 0 U-. 52 50 48 46 0 Oct Nov Dec Jul Aug Sep Oct. Nov 1973 19-74 Pi-gure -.194-7m-YeAm I f 91"@ 1973-7C.i@ m Z@Z -44- 7 @18.4rn Seine --Wing Trawl 6 5 c /e 3 2 1 0 Oct Nov Dec Jul Aug Sep Oct Nov Dec 1973 1974 Figure 19- 1973-74" -45- 112- 78-- 74-- 70-- 66 62- 58- 54, 46-- Oct 73 JuIY'74 Aug:74 Sept'74 Oct 74 Nov '74 CQ Figure f fak&lleqgth'- rf. adl- -46- White Oak River The White Oak River supports a substantial run of blueback herring and smaller runs of alewife, American shad, and hickory shad. Adult blueback herring entered the river in January; they were predominantly four and five year old fish, and most were virgin. Juvenile blueback were found to use the Martin Marietta Belgrade Quarry Lakes, the river below, and certain tributaries as nursery grounds. Their seaward migration was found to occur in October. Adult alewife entered the river in April and were predominantly ages four and five; the vast majority were virgin. Only a few Juvenile American shad were found. All three species of anadromous fish used the Martin Marietta Belgrade Quarry Lakes as spawning areas during April and May. Although the lakes re- present an altered environment, they still appear to support adequate popula- tions of anadromous fish. In addition, in 1975, river herring were found to use Holston and Grant Creeks as spawning areas with spawning probably occurring in Hunter and Webb Creeks. The White Oak River currently supports a small recreational dip net and giii. net iisnery. These appear to have i1ttie or no impact upon the anadr6mous fish populations. The greatest problem facing anadromous fish populations in small rivers is flood control activities such as channelization and watershed development. The impact of such alterations is seen vividly in New River. As a result, it is recommended that future channelization projects be given: closer scrutiny especially where anadromous fish resources are involved. -47- ACKNOWLEDGEIIENTS This project was made possible by the efforts of many of the Division of Marine Fisheries personnel: Walter F. Godwin and Michael W. Street; Fisheries Technicians James J. Bahen, Gerry Davis, and Morris Allison; Lab Technicians Elizabeth Griffin and Dianne Craig; Preston Pate, who assisted in aging scale samples; Secretary Elizabeth Wainwright; and to the many students at Cape Fear Technical Institute who assisted in the field and lab work. Gratitude is expressed to the Martin Marietta Company and its staff at the Belgrade Quarry who provided easy access to the lakes and information. Finally, appreciation is extended to all the commercial and recreational fishermen of the New and White Oak Rivers for their valuable information. -48- LITERATURE CITED Baker, W. D. 1968. A reconnaissance of anadromous fish runs into the inland fishing waters of North Carolina. Final Report of Federal Aid in Fish Restoration, Project AFCS-3. N. C. Wildlife Resources Commission, Raleigh, N. C., 33p. Beal, Kenneth Lee. 1968. Age and growth of the blueback herring (AZosa aestivaZis) (Mitchill). Appendix I. M. S. Thesis, College of William and Mary, Virginia, 82p. Cating, James E. 1953. Determining age of Atlantic shad from their scales. U. S. Fish and Wildlife Service, Fishery Bulletin. 54(85):187-199. Davis, James R. and Randall P. Cheek. 1967. Distribution, food habits, and growth of young clupeids, Cape Fear River system, North Carolina. Proceedings of the 20th Annual Con- ferenc,R of the Southeastern Association of Game and Fish Commissioners, p. 250-260. Davis, James R. and E. G. McCoy. 1965 Survey and classification of the New, White Oak, and Newport Rivers and tributaries 'North Carolina. Final Report for Project F-14-R, Job I-T. North Carolina Wildlife Resources Commission, Raleigh, N.C., 12p. and Appendix. Marcy, Barton C. 1969. Age determinations from scales of (AZosa pseudoharengus) (Wilson), and (AZosa aestivaZis) (Mitchill), in Connecticut wate rs. M. S. Thesis, University of Connecticut, 26p. Nichols, Paul R. and Derrell E. Lauder. 1970 Upstream passage of anadromous fish through navigation locks and use of the stream for spawning and nursery habitat, Cape Fear River, N.C., 1962-1966. U.S. Fish and Wildlife Service, Circular 352, 12p. Stevenson, Charles H. 1899. The shad fisheries of the Atlantic coast of the United States. U. S. Commissioner of Fish and Fisheries, Report for 1898, p. 101-269. Street, Michael W. and Preston P. Pate. 1975. Anadromous fisheries research program, Northern coastal r *egion,* Section I. Final Report for Project AFCS-8. N. C. Division of Marine Fisheries, Morehead City, N. C., 173p. -49- Street, Michael W. and John G. Adams. 1969 Aging of hickory shad and blueback herring in Georgia by the scale method. Georgia Game and Fish Commission', Marine Fisheries Division' Contribution Series No. 18, 13p. Walburg, C. H. and P. R. Nichols. 1967 Biology and management of the*American shad and status of the fisheries, Atlantic Coast of the United States, 1950. U. S. Fish and Wildlife Service, Special Science Report - Fisheries Number 550, 105p. 3 6668 14108 7728