    [From the U.S. Government Printing Office, www.gpo.gov]

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                                                                EFFECTS OF DREDGING AND UNCONFINED DISPOSAL OF
                                                               DREDGED MATERIAL ON MACROBENTHIC COMMUNITIES IN
                                                                               SEWEE BAY, SOUTH CAROLINA'

                                                                                                 by

                                                                                     Robert F. Van Dolah
                                                                                        Dale R. Calder
                                                                                        David M. Knott
                                                                                     Magdalene S. Maclin

                                                                                  Marine Resources Center
                                                          South Carolina Wildlife and Marine Resources Department
                                                                          Charleston, South Carolina 29412

                                                                                                                                                                     00 vs.
                                                                                  Technical Report No. 39                                       091                        1,0 ND
                                                                      South Carolina Marine Resources Center                                      ill                 -     250
                                                                                           April 1979

                                                                       Property of CSC Libraz7

                    Study funded under contract (#DACW-60-77-C-0013) for the U. S. Army Corps of Engineers, Charleston
                   District.
                                                                                                                                                                                   4. t"05-0:-,

                                                                                                                                                            13   1w  -

         Ca
         C"                                             US Department of Commerce
         <=;,                                           NOAA Coactal Services Center Library
                                                        2234 South Hobson Avenue
         CY1.1)
                                                        Charleston, SC                    29405-2413
         =-5

                                                             TABLE OF CONTENTS

                                                                                                                           Page

             LIST OF FIGURES

             LIST OF TABLES

             INTRODUCTION

             DESCRIPTION OF STUDY AREA

             METHODS

                  Benthic Sampling  ....................................................................................       1

                  Benthic Analytical Techniques   .......................................................................      1

                  Water Chemistry  .....................................................................................       3

             RESULTS AND DISCUSSION                                                                                            3

                  Water Chemistry  .....................................................................................       3

                  Benthic Oyster Dredge Collections   ...................................................................      4

                  Benthic Grab Collections  ............................................................................       4

             CONCLUSIONS  ..............................................................................................       7

             RECOMMENDATIONS                                                                                                   8

             ACKNOWLEDGMENTS                                                                                                   8

             LITERATURE CITED  .........................................................................................       9

                                                                     LIST OF FIGURES

                 Figure                                                                                                             Page

                    1. Map of Sewee Bay disposal areas and station locations       ..............................................       2

                    2. Bray-Curtis analysis of Sewee Bay benthic grab collections       .........................................       6

                                                                     LIST OF TABLES

                 Table                                                                                                              Page

                 1.      Hydrographic data collected at subtidal stations in the Sewee Bay area, South Carolina          ...........  10

                 2-14.   Occurrence of epifaunal invertebrates i     n dredge collections from Sewee Bay stations     ..............  12-30

                 15-32.  Macroinvertebrate species collected from the Sewee Bay stations using the Ponar grab          .............  31-51

                    33.  Species diversity, evenness and richness values for Sewee Bay grab samples from August 1977
                         to August-1978   ...................................................................................         52

                    34.  Mean abundance and total number of species collected in the Sewee Bay grab samples from
                         August 1977 to August 1978    .......................................................................        53

                    35.  Wet-weight biomass of Sewee Bay grab samples from August 1977 to August 1978         .....................   54

                                                                             small boats to navigate through the bay. Water
              INTRODUCTION                                                   currents are moderately strong in the area and re-
                                                                             sult from tidal action. Mean tidal range for Sewee
                    Shoaling occurs at a  number of locations in the         Bay is 5.0 feet (National Ocean Survey, 1978).
              Atlantic Intracoastal Waterway (ATWW) through South
              Carolina, and periodic maintenance dredging is                      The disposal sites used in Sewee Bay during
              necessary to keep waterway channels at their pre-              this study were intertidal mud flats adjacent to
              scribed depth. Materials dredged from most shoals              the AIWW but separated from it by several small
              in the waterway through this state are usually                 barrier islands (Fig. 1). This area has infrequent-
              placed in diked disposal areas. However, a shoal               ly received dredged material in the past, with the
              through Sewee Bay, requiring maintenance dredging              last dredging operation occurring in 1964. However,
              in the winter of 1977-1978, has no confined dispo-             no impact studies have been conducted in connection
              sal site nearby. An agreement was made during the              with previous dredging operations in this area.
              annual waterway dredging consultation between the
              Corps of Engineers and several state and federal               METHODS
              agencies in 1977 to undertake unconfined disposal      of
              dredged material in the area.                                  Benthic SampiiR&

                    The impact of dredging and disposal of dredged                Benthic samples were collected at 18 stations
              material in marine and estuarine environments has              in Sewee Bay during the period from August 1977 to
              received increased attention in recent years (for              September 1978. Four of the stations were located
              reviews, see Sherk, 1971; Saila, et al., 1972;                 in the dredged area of the waterway, two in peri-
              Windom, 1976; Morton, 1977), butThe@ reported                  pheral areas near Moores Landing, and 12 in a series
              effects of such activities on benthic communities              of 4 transects at various distances from the dispo-
              are inconsistent. For example, Leathem, et al.                 sal sites (Fig. 1). All stations were sampled prior
              (1973), Slotta, et al. (1973) and Stickney and                 to dredging to provide base line data. Samples were
              Perlmutter (1975TA-served only minimal or short                then collected at every station within two weeks
              term effects from dredging operations while Breuer             after dredging to define the area and extent of
              (1962), Taylor and Saloman (1968), and Kaplan, et              impact. The stations were again sampled three and
              al. (1974) noted more severe and long-term disrup-             six months after dredging to monitor subsequent
              tion of benthic communities resulting from dredging            recovery of the fauna.
              activities. In addition, other studies have noted
              graded effects of dredging and disposal operations                  Qualitative samples were collected at the 13
              on benthic communities which were dependent on dis-            subtidal stations using a modified oyster dredge,
              tance from the disposal sites and depth of the                 towed for three minutes. The dredge used during
              resultant sediment layer (Howell and Shelton, 1970;            this study consisted of a rectangular steel frame
              Saila, et al., 1972; Maurer, et al., 1974; Bingham,            measuring 53 em across the mouth, with a 1 m long
              1978).                                                         bag of 3 cm stretch mesh polypropylene. A skirt of
                                                                             interlacing metal rings protected the bottom of the
                    Due to the variable findings noted above and             bag from chafing. After preliminary sorting of the
              the paucity of dredge-impact studies in areas                  dredge catch on station, unidentified epifaunal
              similar to Sewee Bay, the Division of Marine                   invertebrates and a representative sample of firm
              Resources of the South Carolina Wildlife and                   substrates were preserved in a 10% seawater-formal-
              Marine Resources Department entered into a contract            dehyde solution and returned to the laboratory for
              with the Charleston District of the U. S. Army                 microscopic examination.
              Corps of Engineers to assess the effects of uncon-
              fined disposal of dredged material in this area.                    Quantitative samples were collected at all 18
              The purpose of the study was to determine whether              stations using a 0.05 m2 Ponar grab, Three repli-
              this method could be utilized with minimal distur-             cate samples were taken at each station. After
              bance to the biological communities present and                measuring the volume of each sample, the material
              possibly with less disturbance than would be caused            was placed in large buckets and returned to the
              by diking new areas.                                           laboratory where it was immediately washed through
                                                                             a 1.0 mm sieve. Organisms and sediment remaining
              STUDY AREA                                                     on the sieve after washing were preserved in a 10%
                                                                             seawater-formaldehyde solution and stained with
                    Sewee Bay is a very shallow water mass located           rose bengal to facilitate further sorting under
              west of Bulls Bay about midway along the coast of              magnification. After identification and enumera-
              South Carolina. It is sheltered from the open                  tion, all animals were blotted dry to remove excess
              Atlantic Ocean by Bulls Island. The bay is border-             liquid and weighed on a mettler model PN323 balance
              ed on the west by a series of small islands and                to determine wet-weight biomass.
              bars, created from dredged material left during
              the original dredging of the Atlantic Intracoastal                  Sediment samples were also collected from each
              Waterway through the area. Spartina marshes                    station throughout the study. However, the results
              surround the bay to the south, east, and north.                will not be presented in this report since signifi-
              The AIWW through the Sewee Bay area requires dredg-            cant changes in sediment composition were never
              ing every 10 to 14 years to maintain a channel                 noted (Van Dolah,et al., 1979).
              depth of about 4 m.
                                                                             Benthic Analytical Techniques
                    Although Sewee Bay appears to be relatively
              open at high tide, a substantial portion of its                     Community structure was analyzed using several
              area is exposed as mud flats and oysters bars at               equations from information theory. Species diver-
              low tide. A number of channels, accessible either              sity was measured with Shannon's formula (Pielou,
              from the AIWW or from creeks entering via Bulls                1975):
              Bay to the east, are deep enough (1-3 m) to permit                                     H' = _ZP' 1092 P'

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               where H' is the diversity in bits of information per              224F (APHA, 1971).
               individual, and pi equals-a-1 or the proportion of
                                             N .
               the sample belonging to the ith species. Species                  RESULTS and DISCUSSION
               richness was calculated using the formula:
                                               S-1                               Water Chemistry:
                                        SR = InN
               where S is the number of species and 1nN is the                   Temperature - The shallow waters of Sewee Bay
               natural logarithm of the total number of individuals              rapidly reflected variations in air temperature;
               of all species in the sample. Evenness or equita-                 wide seasonal differences were observed during this
               bility, the distribution of individuals among the                 investigation (Table 1). Temperatures were low
               various species, was measured by:                                 (< 7.0 OC) throughout the study area in January as
                                              H'                                 S7outh Carolina experienced an unusually cold winter.
                                      J' = 1092S                                 Temperatures of approximately 20 'C by late April
               where H' is the species diversity in bits of infor-               and 30 OC by late July were observed in 1978. A
               mation per individual and S is the number of species.             maximum temperature range of 27.3 OC (from a January
                                                                                 reading of 4.7 OC at SB07 to 32.0 OC at SB03 in
                     Faunal homogeneity was measured from both                   July) was noted during the study. Differences from
               qualitative dredge samples and quantitative grab                  station to station were relatively minor on any
               samples. The qualitative measure of similarity                    given sampling date, and differences between surface
               used was the Sorensen coefficient (Sorensen, 1948):               and bottom readings were never large (a maximum of
                                             2C                                  1.8 OC at SBOl in April).      Thus, waters of Sewee
                                      Sl = (a+b)                                 Bay are well-mixed with respect to temperature.
               where a and b are the numbers of species obtained at
               a station on each of the two collection dates and c               Salinity - Although Sewee Bay is influenced to some
               is the number common to both. The quantitative                    extent by the AIWW, it is relatively isolated from
               measure of similarity was the Bray-Curtis coefficient             any major sources of fresh water other than rainfall
               (Boesch, 1977):                                                   and runoff from adjacent land areas. Salinity
                                                                                 measurements taken during the study were all in the
                                       2imin(xii,xii)                            euhaline range (30-40 O/oo), and fluctuations were
                                  S2 = E                                         relatively minor (Table 1). Salinities were highest
                                       i(xij+y-ik)                               in summer and autumn samples and lowest in winter.
               where xij and xik are the number of individuals of                Differences in salinity measurements from surface
               the i th species in two collections under comparison.             to bottom and from station to station on a given
               Using this coefficient, a normal analysis was com-                date were negligible. As Sewee Bay has high and
               pleted on modified data sets with a flexible sorting              stable salinities with no distinct halocline in any
               strategy and standard BETA value of -0.025. All                   direction, it corresponds with the characteristics
               data were log transformed and species which occurred              of a neutral embayment. According to Glooschenko
               in fewer than 4 collections were eliminated from the              and Harriss (1974), a neutral embayment is defined
               analysis. However, prior to this elimination,                     as a partially-enclosed marine environment receiving
               certain species were grouped together to reduce                   negligible fresh water inflow from rivers and with
               error from incomplete identification of damaged                   precipitation approximating evaporation.
               specimens. These groups are as follows: Pinnixa                   Glooschenko and Harriss considered water temperature,
               sayana were lumped with Pinnixa sp.; Panopeus                     light, and nutrients to be the primary factors of
               herbstii with Xanthidae (undet.); Squilla empusa                  ecological importance in temperate neutral embayments.
               with Stomatopoda (undet.); Notomastus lobatus with                Certainly, stresses due to variable salinity on
               Notomastus sp.; Scoloplos          a with Scoloplos sp.;          marine organisms in Sewee Bay are relatively small,
               Haploscoloplos robustus, H. foliosus and H.                       at least for species occurring subtidally. Although
               fragilis with Haploscoloplos sp.; Nemertina sp. A                 it is possible that salinities of the area may be
               with Nemertina (undet.). These groupings make                     reduced somewhat following heavy rainfall, no
               the similarity comparison between collections more                indication of this was evident from the present
               meaningful since, with few exceptions, the specimens              study.
               which could only be partially identified were
               probably the species which could be identified.                   Dissolved Oxygen - Oxygen levels were never observed
                                                                                 to be critically low in Sewee Bay during the study
               Water Chemistry                                                   period (Table 1). Dissolved oxygen values ranged
                                                                                 from a low of 5.0 mg/l in bottom samples from stations
                     Water chemistry was monitored throughout the                SBO8, SB17, and SB18 during summer to a high of
               study using Van Dorn bottles (6-liter capacity) at                10.9 mg/l in surface samples from SB08 and SB10 in
               every station in Sewee Bay except SB02, SB04, SB09,               January. Oxygen readings were lowest in summer,
               SBll and SB16. Samples were collected 1 m below the               highest in winter, and intermediate in spring and
               water surface and 0.3 m above the bottom at each                  autumn. The winter samples, collected within a few
               station. Parameters measured included temperature,                days after dredging and unconfined disposal of
               dissolved oxygen, salinity, turbidity, and total                  dredged material, indicate that the dredging opera-
               and settleable solids. Water temperatures were                    tion was not followed by a widespread and drastic
               measured when the samples were collected using                    reduction in dissolved oxygen levels. Dissolved
               thermometers mounted in the Van Dorn bottles.                     oxygen levels could not be recorded for samples
               Dissolved oxygen and turbidity samples were fixed                 taken immediately after dredging at station SB04 due
               immediately after collection. In the laboratory,                  to very high silt levels. However, the epibenthic
               dissolved oxygen was analyzed by the modified                     sample collected shortly thereafter smelled noxious,
               Winkler titration method. Turbidity and salinity                  which may have been attributable either to anaerobic
               were measured using a Hach Model 2100A Turbidi-                   conditions, or smothering and subsequent decomposi-
               meter and a Beckman Model RS7B Induction Salino-                  tion of some bottom-dwelling organisms.
               meter. Solids were analyzed by American Public
               Health Association (APHA) Standard Methods 224C and                     The low oxygen values observed in summer are due

                                                                             3

                 to a combination of factors, including high water              Oyster dredge samples were taken at four
                 temperatures and the respiration of organisms.            stations (SB03, SB07, SB12, SB17) along the axis
                 However, D.O. levels never departed significantly         of Sewee Bay (Fig. 1). Of these, SB03 was
                 from saturation levels. The effects of dredging           especially noteworthy because it had the richest
                 would probably have been much more adverse had the        assemblage of epifaunal species found anywhere
                 operation been conducted at any other season of           in Sewee Bay during the study. The bottom at this
                 the year, but especially in summer when oxygen            station consisted of relatively mud-free shell,
                 levels are low and community metabolism is high.          and an average of 43 species was collected over
                 Waterway dredging through Sewee Bay was fortuitously      the four sampling periods. While the minimum
                 well-timed to coincide with a period when its             number of species collected (32) came immediately
                 environmental impact, particularly with respect           after dredging, there was no evidence that the
                 to D.O., was probably lowest.                             drop in species numbers could be attributed to
                                                                           smothering from spreading dredged material.
                 Turbidity and Solids - Values for these parameters        Instead, many of the epifaunal species found there,
                 were rather high throughout the year in Sewee Bay.        especially certain hydroids, entoprocts, and
                 Pronounced increases as a result of dredging were         sponges, undergo a period of dormancy in winter
                 apparent in winter samples from four stations             due to low water temperatures and were simply not
                 (SBOI, SB03, SB04, SB05), while samples from the          active during February. Species numbers and
                 remaining stations at this time were generally            community composition in spring and summer samples
                 comparable to, or even lower than, those collected        were very similar to those noted in the pre-dredge
                 during other seasons of the year (Table 1). It is         survey (Table 3). The bottom at station SB07
                 particularly noteworthy that samples from stations        consisted largely of broken shell and mud, but a
                 SBO1, SB03, SB04, and SB05 were taken within 4-5          moderately diverse faunal assemblage was present
                 days after dredging and unconfined disposal of            (Table 6). No adverse effects were apparent from
                 dredged material in and adjacent to these sites,          the disposal of dredged material nearby. Two other
                 and while the dredge was still operating in the           stations in the middle of Sewee Bay (SB12, SB17)
                 waterway through Sewee Bay. Samples from the              were located in muddy, shallow water areas. The
                 remaining stations were collected shortly after           epifauna at both stations, and particularly SB12,
                 all dredging in the area had been completed. This         was impoverished and no changes were detectable
                 indicates that abnormally high levels of turbidity        that were obviously related to disposal practices
                 and solids resulting from the dredging operations         (Tables 9, 13).
                 were of very short duration. Several of the
                 turbidity and solids values reached their lowest               Four stations were also sampled at the mouths
                 levels in winter. This corresponds with the data          of creeks along the eastern edge of Sewee Bay
                 of Mathews and Shealy (1978), who observed that           (Fig. 1). The relatively low number of species
                 turbidities and solids in the Edisto and Cooper           observed in February samples at stations SB08, SB13,
                 estuaries were lower in winter than at any other          and SB18 was again attributed largely to the
                 time of the year.                                         seasonal dormancy of a number of sponges, hydroids,
                                                                           entoprocts, and others (Tables 7, 10, 14).
                 Benthic Oyster Dredge Collections:                        However, the reduction in species numbers collected
                                                                           at station SB04 was drastic (Table 4), and clear
                     A moderately diverse assemblage of species was        evidence of smothering by sediments from unconfined
                 noted in the epifauna of the Sewee Bay area. Over-        disposal of dredged material was apparent at the
                 all, species composition was relatively similar           time of sampling. A considerable amount of soft
                 from one location to another, especially before           mud was taken in the February oyster dredge tow.
                 dredging operations were undertaken (Tables 2-14).        In addition, dead mud crabs and bivalves (Tagelus
                 For example, a suite of relatively stenohaline            plebeius, Tellina sp.) were collected and the
                 bryozoans (Bugula neritina, Schizoporella errata,         entire sample was malodorous. No appreciable
                 Hippoporina verrilli, Microporella ciliata,               recovery of the fauna was observed when sampling
                 Parasmittina--nitida) was present at the majority         was conducted in late April. By the summer of 1978,
                 of the 13 stations where sampling with a modified         the community was still less diverse than before
                 oyster dredge was conducted.                              dredging but a substantial increase in the number
                                                                           of species had occurred over the winter and spring
                     Physical disruption of epibenthic assemblages,        samples, and faunal homogeneity had increased
                 caused by dredging of the Atlantic Intracoastal           markedly (Table 4).
                 Waterway during January of 1978, was clearly
                 evident in samples from stations SBOl. SB05, SBIO,             Finally, oyster dredge collections were taken
                 and SB15 (Tables 2, 5, 8, 12). The average number         at station SB14 in a small creek west of the AIWW.
                 of species at these four waterway stations was 28         This station was relatively isolated from disposal
                 before dredging and 6 immediately after. Repopu-          sites and no impact was observed on the epifauna
                 lation was noted'in subsequent sampling intervals.        of the area (Table 11).
                 Three months after dredging, the average number of
                 species at the four stations in the AIWW had risen        Benthic Grab Collections:
                 to 20. Six months after dredging, the average
                 number of species was 22, only slightly lower than        Species Composition - A total of 129 identifiable
                 the number observed the previous summer. Faunal           species was collected by the Ponar grab in Sewee
                 homogeneity between pre-dredge and post-dredge            Bay (Tables 15-32). Of these, four polychaete
                 collections was low to moderately low immediately         species, Lumbrineris tenuis, Paraprionospio pinnata,
                 after dredging but increased substantially by the         Heteromastus filiformis, and Nereis succinea,
                 end of the study (Tables 2, 5, 8, 12). The total          comprised over 46% of the total number of animals
                 catch of animals and hard substrate was usually           collected throughout the study period. These
                 small both before and after dredging at all of the        species, as well as several others, were quite
                 stations in the waterway.                                 ubiquitous in their distribution among stations.

                                                                      4

           For example, L. tenuis, H. filiformis and N. succinea       of dredged material (Tables 20, 25, 30). All of these
           were present at all 18 stations at least once during        stations were separated from the disposal sites by
           the study period while P. pinnata was observed at 17        short projections of land which may have deflected much
           stations during the one-year period.                        of the dredged sediments. Tn addition, many of the
                                                                       species at these stations have been categorized as
                The community structure of the channel stations        pollution resistant (Wass, 1967; Saila, et al., 1972:
           (SBOl, SB05, SB10, SB15) was dominated by P. pinnata        Maurer, et al., 1978) and could have surviv7ed the
           in the late summer prior to dredging. Oth@er abun-          deposition which did occur. The Bray-Curtis coefficient
           dant species generally present at this time included        analysis of these stations indicated that the species
           the decapod Ogyrides limicola and two polychaetes,          composition was similar throughout the year at SB11,
           L. tenuis (all stations) and Glycinde solitaria             and similar during the first three sampling quarters at
           (SId5). Following the winter dredging operation,            SB06 (Fig. 2). The decline in community complexity and
           P. pinnata densities had decreased at all stations          density during the fourth quarter sampling period at
           and, at two stations, it was replaced in terms of           SB06 (Table 20) was probably not due to disposal effects
           numerical dominance by the pelecypod Mulinia                as similar decreases were not observed in the third
           lateralis (SB05) and the decapod 0. limicola                quarter sampling effort. At station SB16, community
           (SB10).                                                     structure was dissimilar throughout most of the study
                                                                       period (Fig. 2). However, it is unlikely that dredged
                An analysis of overall species composition             material had much influence as a strong tidal current
           using the Bray-Curtis similarity coefficient showed         was present and spoil deposits were never observed.
           no consistent or distinctive patterns among channel
           stations with respect to dredge effects. Six months              The communities at stations located in the middle
           after dredging, species composition was only slightly       of Sewee Bay (SB03, SB07, SB12, SB17) were not severely
           similar to the pre-dredge species composition at            affected by spoil deposition. During the initial
           stations SBOI and SB05, and dissimilar at stations          sampling period (August, 1977), the community at SB03
           SB10 and SB15 (Fig. 2) even though these sampling           was depauperate and dominated by L. tenuis. After
           periods were in equivalent seasons. However,                spoil deposition, the community sampled was much more
           comparable trends were observed at stations which           complex (Table 17) and remained diverse throughout the
           should not have been affected by the dredging               rest of the study period, although species composition
           operation (discussed later). Thus, some of the              was dissimilar on each sampling date (Fig. 2). Para-
           changes in community structure may have been due            prionospio pinnata, which was present at the last three
           to normal seasonal and sampling variations.                 sampling quarters, may have been introduced to the
                                                                       community through deposition as it was not present in
                Alterations in species composition which               the first quarter. Stations SB07, SB12 and SB17 showed
           might be attributable to the dredging operation             trends similar to SB03 although initial community
           were noted at station SBOl. Three months after              structure was more complex. Spoil deposits were present
           dredging, there was a substantial increase in               at SB17 in the second and third sampling quarters and
           the number of species normally associated with              may account for some of the changes observed (Fig. 2).
           "live" bottom communities at this station (Table            However, sediment characteristics were not visibly
           15). Therefore, it is possible that the dredge              altered at SB07 and SB12, and community structure was
           removed soft mud from the channel down to a firmer          different during equivalent seasons at SB12 but similar
           substrate which permitted a "live" bottom community         at SB07 (Fig. 2). Therefore, it is likely that much of
           to form. However, this community is either patchy           the variation in species composition at all of these
           or was smothered by channel wall slump since a              stations is due to a combination of seasonal effects
           similar species composition was not observed during         and sampling variability resulting from patchy community
           the fourth quarter sampling period.                         distribution.

                Community structure at stations nearest the                 Stations farthest from the disposal sites (SB04,
           disposal sites exhibited varied effects from                SB08, SB13, and SB18) also showed changes in community
           deposition of dredged material.   Station SB02              structure immediately after, and three months after,
           was situated very close to a disposal site and              deposition of dredged material. Soft, oozy sediments
           was the most severely affected. Prior to dredging,          were present at stations SB04, SB08, and SB18 on these
           the community at this station was dominated by the          sampling dates and probably accounted for some of the
           polychaetes L. tenuis and Aricidea fragilis                 changes observed. For example, P. pinnata was a
           (Table 16). Dwr-ing the se_condquar@tersampling             dominant species at two of these stations immediately
           period, the dominant species was H. filiformis,             after deposition, although it was not present before.
           but L. tenuis remained an important component of            As this species was common in the channel prior to
           the community. Most of the changes which occurred           dredging, it may have been introduced with the spoil.
           at this time were probably seasonal,as dredged              However, it is unlikely that spoil disposal severely
           material was still being deposited nearby. Three            influenced the communities through smothering since a
           months after spoil deposition, however, the                 diverse assemblage of species was always present. Six
           community was reduced to one infaunal species,              months after dredging, the species composition at these
           N. succinea (a spoil-resistant polychaete;                  stations was slightly similar to pre-dredge species
           Iffaurer, et al., 1978) and two epifaunal crustacean        composition (Fig. 2). The community structure at SB13,
           species (each represented by one individual).               on the other hand, was dissimilar on every sampling
           Thus, the original community was smothered and              date (Fig. 2). Even so, no decline in community
           although community complexity had increased to   a          complexity at SB13 occurred as a result of spoil
           pre-dredge level six months after dredged material          deposition.
           deposition, the species composition was altered
           (Table 16) and dissimilar by Bray-Curtis                        Two stations which were not influenced by the
           similarity coefficient analysis (Fig. 2).                   dredging operation were SB09 and SB14. Both of these
                                                                       stations were out of the channel and on the opposite side
                Stations SB06, SB11, and SB16, on the other            of the waterway with respect to the disposal sites. Thus,
           hand, were not noticeably affected by deposition            changes which occurred at these stations were most likely

                                                                     5

                            Figure 2.           Bray-Curtis analysis of the Sewee Bay benthic grab collections.
                                                First Quarter = samples collected before dredging; Second Quarter
                                                samples collected immediately after dredging; Third Quarter
                                                samples collected 3 months after dredging; Fourth Quarter = samples
                                                collected 6 months after dredging and 1 year after first quarter
                                                samples.                                            SIMILARITY
                                                                     .8         .6          .4         .2          0          -.2        -.4          -.6

                                                       STATION(OUMTER)
                                                                 1(1)
                                                                 :
                                                                 00)
                                                                 5 (1)
                                                                 90)
                                                                 140)
                                                                 13(4)
                                                    A            1? (2)
                                                                 40)
                                                                 3(4)
                                                                 50)
                                                                 1514) - - - - - - -
                                                                 4
                                                                 OW
                                                    B            1 (4)
                                                                 5(4)
                                                                 10(2)

                                                                 5
                                                    C            5(3)
                                                  - - _13
                                                    D            I W14
                                                           --I 5L    - - - - - - - - -
                                                                 2. ?41
                                                                 4(2)
                                                    E            6(4)
                                                            _17,it4) - - - - - - - - -
                                                                 14(3)
                                                                 13(3)
                                                                 18(3)
                                                                 4(3)
                                                                 170)
                                                                 3(3)
                                                                 8(3)-
                                                                 16(3)
                                                    F            6(2)
                                                                 9(2)
                                                                 12(3)
                                                                 9(3)
                                                                 100)
                                                                 18(2)
                                                                 8(2)
                                                                 ? (2)
                                                                 1(3)

                                                                 170)
                                                    G            7(4)
                                                                 18(4)
                                                                 30  - - - - - - - - -
                                             - - - -             14-T4)
                                                                 2(2)
                                                                 12(1)
                                                                 6(3)
                                                                 60)
                                                    H            16(l)
                                                                 G(2)
                                                                 30)
                                                                 130)
                                                                 20)
                                                                 9A
                                                           -19 )                                  -
                                                                 IU(4) - - - - - -
                                                                 11(3)
                                                                 1(1)

                                                                 1000
                                                                 12(,Q
                                                                 110
                                                                     9@=7

                                             - - - -             12-0
                                                                 2(3)-

                                                                     .8         .6          .4         .2           0         -2          -.4         -.6
                                                                                                    SIMILARITY
                                                                                                   6

              due to seasonal effects. Station SB09 was domin-              the rest of the study period. The observed increase
              ated by L. tenuis throughout the study period                 was the result of increases in both evenness and
              except in January (second quarter) when M.                    species richness. H' values fluctuated at stations
              lateralis was the dominant species (Table 23).                SB04 and SB08 and had declined to relatively low
              Sporadic increases in M. lateralis densities                  values by the fourth qjjarter sampling period due to
              at this time of the year have also been noted                 decreased evenness and species richness. H' values
              in previous studies (Boesch, 1973; Stickney and               at SB09 remained similar throughout this study
              Perlmutter, 1975). The community structure at                 although species richness did increase during the
              SB14 was variable, partly due to seasonal effects.            winter and spring months.
              However, this station had an extremely hard bottom
              with strong currents present, and much of the                 Abundance - An analysis of the mean abundance of
              variability is attributable to poor grab samples.             the benthic communities at each station (Table 34)
              At both stations, the community was similar in the            indicated that the dredging operation had some
              second and third quarters but dissimilar in the               short term, localized effects in Sewee Bay. As
              first and fourth quarters (Fig. 2).                           expected, mean densities at all channel stations
                                                                            (SBO1, SB05, SB10, SB15) declined immediately after
              Species Diversity -The analysis of species                    dredging, whereas densities had increased during
              diversity (H'), evenness (J') and species richness            this period at most of the stations farthest from
              (SR) of benthic communities in Sewee Bay did not              the dredging activity (SB04, SBO8, SB09, SB14, SB18).
              reflect any widespread changes resulting from the             The percent of decrease at the channel stations
              dredging operation (Table 33).                                varied, ranging from a minimum of 49% at SB10 to a
                                                                            maximum of 85.2% at SB15. However, declines in
                    H' values increased immediately after dredg-            abundance were only significant (P<0.05 ANOVA test)
              ing at two of the channel stations (SB05, SB15)               at three of the channel stations (SB05, SB10, SB15)
              and all channel stations had higher H' values                 and, at all but one station (SB15), abundance had
              three months after the dredge had passed through              increased to pre-dredge levels by April.
              the area. This increase was primarily due to
              increased species richness and evenness at                         Significant density changes (P<.05, ANOVA test)
              stations SBO1, SB10 and SB15 and increased                    were also observed at two other stations. The
              evenness at station SB05. Six months after                    community at station SB16 increased during the
              dredging, H' had decreased at all channel                     second quarter sampling period, and the community
              stations to values which were approximately                   at station SB17 decreased in abundance during the
              equivalent to those observed in the first quarter             fourth quarter sampling period. These changes were
              sampling period.                                              probably due to seasonal effects or sampling varia-
                                                                            bility as discussed previously. Fluctuations in
                    Species diversity patterns were more varied             mean densities at all other stations were not signi-
              at stations nearest the disposal sites. The                   ficant during the study period (P>.05, ANOVA test).
              communities at all of these stations (SB02, SB06,
              SB11, SB16) were more diverse immediately after               Biomass - Wet-weight biomass estimates (Table 35)
              dredged material disposal when compared with the              varied from a low of 0.06 gms at stations SB02 and
              pre-dredge, August sampling period. H' values                 SB03 (August 1977, 1978) to a high of 7.02 gms at
              continued to increase at station SBll and SB16                station SB15 (August, 1977) with no significant
              during the third quarter sampling period before               patterns discernable. The observed variability is
              declining to values approaching those observed                due to (1) the low biomass present at these stations
              initially. H' values at station SB06 remained                 on any sampling date, (2) high variability between
              the same throughout the year with the exception               replicate samples, and (3) the occasional presence
              of the increase noted in January. Diversity                   of large bivalves, gastropods, anemones, etc. in the
              values for station SB02, on the other hand,                   samples. The effects of these factors are best
              reflected the demise of this community through                exemplified by an analysis of two stations. Pre-
              smothering as H', V and SR values were quite                  vious data indicated that SB02 was the most severely
              low three months after spoil deposition.                      affected station in Sewee Bay through smothering
              However, by the end of the study period, all                  from deposition of dredged material which resulted
              three indices had increased at station SB02 to                in a substantial decline in community complexity and
              values higher than those observed prior to the                diversity by April, 1978. However, by April biomass
              dredging operation.                                           had increased to the highest values observed due to
                                                                            the presence of several relatively large N. succinea.
                    Species diversity (H') was consistently                 The community at station SB14, on the ot        band,
              higher after spoil deposition at all of the                   was not significantly influenced by the dredging
              stations located in the middle of Sewee Bay                   operation because of its location. Nonetheless,
              (SB03, SB07, Shl2) except station SB17. This                  biomass decreased from a high initial value of 2.89
              was primarily due to increased species richness.              gms in August, 1977 to a low of 0.27 gms by April,
              At station SB17, H' values also increased in                  1978. Thus, it is difficult to interpret changes
              January but later declined to very low values                 in biomass with respect to effects of dredging and
              by the end of this study. These values are                    disposal of dredged material.
              probably an artifact of sampling variability,
              however, as similar values should have been
              observed in April if deposition of dredged                    CONCLUSIONS
              material was the cause for this decline.                           In summary, the effects of dredging and uncon-
                                                                            fined disposal of dredged material on epifaunal
                    Diversity values varied at the stations                 macroinvertebrate communities were detectable at
              farthest from the disposal sites. H' increased                five of the 13 stations sampled with an oyster dredge
              at stations SB13, SB14 and SB18 after the dredg-              in Sewee Bay. Disruption of the epibenthos by
              ing operation, and remained higher throughout                 channel dredging was apparent at each of the four

                                                                        7

                  stations located in the AIWW. However, the epi-              the area.
                  fauna was not especially rich at any of these four
                  stations before dredging, and faunal composition                  2) Good flushing patterns present at many Sewee
                  at the end of the study approached that observed             Bay stations, as well as flow diversion by land and
                  during the baseline survey conducted in the summer           oyster hummocks which protected other areas where
                  of 1977. Of the eight stations in Sewee Bay proper           stations were located.
                  where dredge tows were made, a severe impact on the
                  benthos was noted only at SB04. Evidently, dredged                3) The presence of species at several stations
                  material from the disposal area was carried by               that are resistant to spoil deposition or unhealthy
                  water currents into the creek where SB04 was located.        conditions.
                  Some recovery was noted at this station six months
                  after dredging. Species numbers were generally                    4) The presence of several ubiquitous species
                  lower at all stations in the winter after dredging           in Sewee Bay, permitting rapid recolonization in
                  when compared with samples collected during the              areas which were affected.
                  summer. This was attributed largely to normal
                  seasonal differences in community composition. The                5) The similarity of sediment composition
                  wide annual range of water temperature influenced            present in the channel and disposal areas of Sewee
                  the activity patterns of several epifaunal species.          Bay.
                  A number of sponges, hydroids, entoprocts, and
                  ascidians were observed in a dormant state during                 6) Seasonal effects, which minimized adverse
                  the winter sampling period. These species returned           water quality conditions during the winter dredging
                  to activity with the return of warmer water tempera-         period, and caused increased community diversity at
                  tures in spring.                                             most stations afterwards; thereby countering dredge
                                                                               and disposal effects.
                       The effects  of dredging on the infaunal commun-
                  ities in Sewee Bay were short term and localized.            RECOMMENDATIONS
                  Immediately after dredging, the community structure
                  at most channel stations was different when compared              Disposal of dredged material can be managed to
                  with pre-dredge samples. Total animal abundance and          minimize adverse effects in the estuarine environment.
                  biomass had also decreased. Although some of these           Depending upon the timing and frequency of disturbance,
                  changes are due to seasonal effects, dredging                Rhoads, et al. (1978) suggested that the productivity
                  obviously affected the channel communities through           of benttd-c -communities could actually be enhanced by
                  bottom disturbance. However, this disruption did             dredge-spoil disposal operations. Although the
                  not adversely affect species diversity which in-             following recommendations may not enhance benthic
                  creased after dredging due to a decline in P. pinnata        productivity, adverse effects should be minimized
                  dominance, and a concomitant increase in species             in Sewee Bay if:
                  richness. Six months after the dredge had passed
                  .through the area, all stations showed indications of             1) Dredging and disposal of dredged material
                  returning to pre-dredge conditions. The lack of any          are undertaken in late fall and/or early winter to
                  long term, adverse effects in the channel correlates         reduce adverse effects on recruitment patterns.
                  well with studies on the intracoastal waterway in
                  Georgia (Stickney and Perlmutter, 1975) and in                    2) Dredging operations are conducted relatively
                  Delaware Bay (Maurer, et al., 1974).                         often, so that the volume of dredged material is not
                                                                               large at any given time.
                       The effects of unconfined disposal on the in-
                  fauna in Sewee Bay were varied and somewhat depen-                3) The practice of using several disposal sites
                  dent on distance from the points of deposition.              is continued in any future operations.
                  The intertidal station, SB02, was the most severely
                  affected through smothering which altered community          ACKNOWLEDGMENTS
                  structure and decreased species diversity, but did
                  not affect community biomass and numerical abundance.             We are greatly indebted to a number of our co-
                  Other stations in close proximity to the disposal            workers who assisted us during the study. Mary Jo
                  sites did not appear to be adversely affected by             Clise, Rachel Goldberg, Paula Keener, Priscilla
                  deposition. The community composition at stations            Knight, Parker Lumpkin, Kathleen Meuli, Todd Nimmich,
                  further from the spoil sites changed during the              Rick Richter, and James Scoggin were particularly
                  study period, but many changes observed were due             helpful in various aspects of the field work.
                  to normal seasonal variability.                              Thanks are due to W. Z. Carson, Rob Dunlap, and Don
                                                                               Marchette for allowing us to use their boats.
                       In conclusion, the influence of dredging                Water chemistry samples were analyzed by Kathy
                  operations on benthic communities in Sewee Bay were          Austin and Virginia Young.    Dr. E. L. Bousfield
                  short term and isolated. Furthermore, it is un-              verified the identification of several amphipod
                  likely that dredged material disposal had a profound         species in our samples. We    wish to thank John
                  effect at any station in terms of the ecological             Carothers, Robbin Blackman,   and Steve Morrison of
                  interactions of benthic communities with higher              the Charleston District, U.   S. Army Corps of
                  trophiclevels. These results differ from other               Engineers, for their part in the study design and
                  studies in which larger areas were noticeably                for their encouragement and support of our work.
                  smothered and subsequently recolonized (Saila, et            Chris Brosseau, Nicky Kopacka, and Lourene Rigsbee
                  al., 1972; Maurer, et al., 1974; Bingham, 1978).             assisted in the processing of data. Mary Anne
                  T-he absence of widespread, long-term effects in this        Carson, Pat DuPree, and Lexa Ford typed the manu-
                  study may be due to several factors. These include:          script. M. H. Shealy, Jr. critically reviewed the
                                                                               manuscript. This study was supported under Contract
                       1) The use of several disposal sites in Sewee           Number DACW60-77-C-0013 from the U. S. Army Corps
                  Bay which spread dredged material more evenly over           of Engineers, Charleston District.

                                                                          8

                                                                               Maurer, D. L., R. T. Keck, J. C. Tinsman, W. A.
              LITERATURE CITED                                                       Tinsman, W. A. Leathem, C. A. Wethe, M.
                                                                                     Huntzinger, C. Lord, T. M. Church. 1978.
              American Public Health Association. 1971. Standard                     Vertical migration of benthos in simulated
                    methods for the examination of water and waste-                  dredged material overburdens. Vol. 1: Marine
                    water including bottom sediments and sludges.                    benthos. U. S. Army Engineer Waterways
                    13th ed. Amer. Public Health Assoc. Inc., New                    Experiment Station, Tech. Rept. D-78-35. 97 p.
                    York, N. Y. 769 p.
                                                                               Morton, J. W. 1977. Ecological effects of dredging
              Bingham, C. R. 1978. Aquatic disposal field inves-                     and dredge spoil disposal: A literature review.
                    tigations Duwamish waterway disposal site Puget                  Technical Paper #94, U. S. Fish and Wildlife
                    Sound, Washington. Appendix G: Benthic                           Service. 33 p.
                    community structural changes resulting from
                    dredged material disposal Elliott Bay disposal             Pielou, E. C. 1975. Ecological diversity. Wiley-
                    site. U. S. Army Engineer Waterways Experiment                   Interscience, New York. 165 p.
                    Station, Tech. Rept. D-77-24. 103 p.
                                                                               Rhoads, D. C., P. L. McCall, and J. Y. Yingst.
              Boesch, D. F. 1973. Classification and community                       1978. Distribution and production on the
                    structure of macrobenthos in the Hampton                         estuarine sea floor. Amer. Sci. 66: 577-586.
                    Rhoads area, Virginia.Mar. Biol. 21: 226-244.
              Boesch, D. F. 1977.       Application of numerical               Saila, S. B., S. D. Pratt, and T. T. Polgar. 1972.
                    classification in ecological investigations                      Dredge spoil disposal in Rhode Island Sound.
                    of water pollution. Virginia Inst. Mar. Sci.                     Univ. Rhode Island. Mar. Tech. Rep. 2.
                    Spec.  Sci. Rep. No. 77. 113 p.                                  48 pp.
              Breuer, J.   P. 1962. An ecological survey of the                Sherk, J. A., Jr. 1971. The effects of suspended
                    lower  Laguna Madre of Texas, 1953-1959. Publ.                   and deposited sediments on estuarine organisms:
                    Inst.  Mar. Sci., Univ. Tex. 8: 153-185.                         Literature summary and research needs. Chesa-
                                                                                     peake Biol. Lab., Solofnons, Md., Contrib. No.
              Calder, D.   R., B. B. Boothe, Jr., and M. S. Maclin.                  443. 73 pp.
                    1977. A preliminary report on estuarine                    Slotta, L. S., C. K. Sollitt, D. A. Bella, D. R.
                    macrobenthos of the Edisto and Santee River                      Hancock, J. E. McCauley, and R. Parr. 1973.
                    systems, South Carolina. South Carolina Mar.                     Effects of hopper dredging and in channel
                    Res. Center Tech. Rep. No. 22. 50 p.                             spoiling in Coos Bay, Oregon. Oregon State
              Glooschenko, W. A., and R. C. Harriss. 1974.                           Univ., Corvallis. 141 pp.
                    Neutral embayments. Pages 488-497. In                      Sorensen, T. A. 1978. A method of establishing
                    H. T. Odum, B. J. Copeland, and E. A. McMahan                    groups of equal amplitude in plant sociology
                    (eds.), Coastal ecological systems of the                        based on similarity of species content and its
                    United States, Vol. I. The Conservation                          application to analyses of the vegetation on
                    Foundation, Washington, D. C.                                    Danis commons. K. dan5ke Vidensk. Selsk. Biol.
              Howell, B. R. and R. G. J. Shelton. 1970. The                          Skr. 5(4): 1-34.
                    effect of china clay on the bottom fauna of                Stickney, R. R., and D. Perlmutter. 1975. Impact
                    St. Amstell and Mevagissy Bays. Jour. Mar.                       of intracoastal waterway maintenance dredging
                    Biol. Ass. U.K. 50: 593-607.                                     on a mud bottom bentho5 community. Biol.
              Kaplan, E. H., J. R. Welker, and M. G. Kraus.                          Conserv. 7: 211-226.
                    1974. Some effects of dredging on popula-                  Taylor, J. L., and C. H. Saloman. 1968. Some
                    tions of macrobenthic organisms. U. S.                           effects of hydraulic dredging and coastal
                    Natl. Mar. Fish. Serv., Fish. Bull.                              development in Boca Ciaga Bay, Florida. U. S.
                    72(2): 445-480.                                                  Fish Wildl. Serv. Fish. Bull. 67(2): 213-241.
              Leathem, W., P. Kinner, D. Maurer, R. Biggs,                     Van Dolah, R. F., D. R. Calder, F. W. Stapor, Jr.,
                    and W. Treasure. 1973. Effects of spoil                          R. H. Dunlap, and C. R. Richter. 1979.
                    disposal on benthic invertebrates. Mar.                          Atlantic intracoastal waterway environmental
                    Pollut. Bull. 4(8)* 122-125.                                     studies at Sewee Bay and North Edisto River.
              Mathews, T. D., and M. H. Shealy, Jr. 1978.                            Final Report, Contract #DACW-60-77-C-0013,
                    Hydrography of South Carolina estuaries,                         Charleston District, U. S. Army, Corps of
                    with emphasis on the North and South Edisto                      Engineers. 146 p.
                    and Cooper Rivers. South Carolina Mar. Res.                Wass, M. L. 1967. Biological and physiological
                    Center Tech. Rep. No. 30. 148 p.                                 basis of indicator organisms and communities:
                                                                                     In: Olson, T. A. and F. J. Burgess, Pollution
              Maurer, D., R. Biggs, W. Leathem, P. Kinner,                           _i@n_d Marine Ecology, Interscience, N. Y. pp. 271-
                    W. Treasure, M. Otley, L. Watling, and                           283.
                    V. Klemas. 1974. Effect of spoil dis-
                    posal on benthic communities near the mouth                Windom, H. L.   1976.   Environmental aspects of
                    of Delaware Bay.    College of Marine Studies,                   dredging in the coastal zone.     CRC Critical
                    Univ. of Delaware, Lewes and Newark.         231 pp.             Rev. in Environ. Control.     7: 91-109.

                                                                           9

             Table 1. Hydrographic data collected at subtidal stations in the Sewee Bay area, South Carolina.

                                                                                                                                    SOLIDS
                                         STATION                     TEMP.      SALINITY        D.O.       TURBIDITY                      (mg/1)
             STATION         DATE         DEPTH          DEPTH        (C)         ('/oo)       (mg/1)         (FTU)        TOTAL      SETTLEABLE

              SBOl        10-19-77           4         Surface       17.1         34.53          7.1          16.0         110.4          38.8
                                                       Bottom        16.9         34.58          7.4          19.0         110.4          80.8
                            1-17-78                    Surface        7.0         30.62          9.2          14.0            -              -
                                                       Bottom         6.9         31.89          9.4          .68.0        189.2          39.2
                            4-24-78                    Surface       20.9         32.94          7.3           9.0          96.8          13.2
                                                       Bottom        19.1         32.94          6.9          26.0         165.6          28.0
                            7-24-78                    Surface       30.2         34.10          5.6          12.0         103.2          76.4
                                                       Bottom        30.1         34.15          5.8          17.0         137.2          75.6
              SB03        1110-18-77         1         Surface       16.6         34.66          7.3          12.0         104.4          28.0
                                                       Bottom        16.5         34.73          7.6          12.0         126.8          28.4
                            1-17-78                    Surface        6.9         32.01          9.4          30.0         131.2          30.4
                                                       Bottom         6.9         32.09          9.5          44.0         179.2          80.0
                            4-24-78                    Surface       19.0         32.98          7.0          21.0            -              -
                                                       Bottom        19.0         32.89          6.9          19.0            -              -
                            7-24-78                    Surface       32.0         34.07          5.5          16.0         120.4          31.6
                                                       Bottom        32.0         34.07          5.6          18.0         138.0          41.6

              SB04        10-18-77           3         Surface       16.8         34.69          7.7          11.0         101.2          30.4
                                                       Bottom        16.8         34.74          7.7          34.0         106.4          24.8
                            1-17-78                    Surface        6.5         32.07          9.6          34.0         134.4          56.0
                                                       Bottom         6.4
                            4-24-78                    Surface       20.1         32.98          7.2          12.0         103.6          10.4
                                                       Bottom        19.8         33.00          7.2          16.0         131.6          14.4
                            7-24-78                    Surface       29.9         34.00          5.4          20.0         127.6          60.8
                                                       Bottom        29.9         34.02          5.2          21.0          86.8          21.2

              SB05        10-19-77           4         Surface       17.4         34.31          7.2          12.0         115.2          30.4
                                                       Bottom        16.9         35.48          7.2          17.0            -              -
                            1-17-78                    Surface        6.9         32.16          9.7          23.0         154.8          58.8
                                                       Bottom         6.7         32.09          9.7          45.0            -              -
                            4-24-78                    Surface       20.3         32.99          7.2            7.5           -              -
                                                       Bottom        20.1         33.00          6.9          15.0            -              -
                            7-24-78                    Surface       29.8         34.31          5.6          11.0            -              -
                                                       Bottom        29.5         34.31          5.2          18.0         121.2          30.0

              SB07        10-18-77           2         Surface       16.6         34.61          7.5           8.9         108.8          36.0
                                                       Bottom        16.5         34.58          7.5          11.0         121.6          40.8
                            2-1-78                     Surface        4.8         30.57         10.7          16.0          82.0           2.4
                                                       Bottom         4.7         30.62           -           17.0          77.0           1.8
                            4-24-78                    Surface       19.7         32.97          6.9          13.0         116.0          19.2
                                                       Bottom        19.7         33.02          6.9          13.0         135.6          10.4
                            7-24-78                    Surface       29.8         34.26          5.5          15.0         138.4          19.6
                                                       Bottom        29.8         34.30          5.5          19.0         107.6          29.6

              SB08        10-18-77           2         Surface       16.5         34.61          7.6          14.0         117.2          21.2
                                                       Bottom        16.5         34.61          7.8          15.0         134.4           9.2
                            2-1-78                     Surface        4.9         30.62         10.9          18.0          88.0          25.2
                                                       Bottom         5.0         30.60         10.7          17.0          96.0          34.4
                            4-24-78                    Surface       20.0         32.90          6.9          17.0            -              -
                                                       Bottom        20.0         33.01          6.9          19.0         160.0          27.6
                            7-24-78                    Surface       30.0         34.03          5.1          21.0         150.0          43.2
                                                       Bottom        30.0         34.05          5.0          24.0         147.2           2.0

              SB10          8-22-77          4         Surface       27.4         33.66          6.4          12.0                           -
                                                       Bottom        27.0         33.62          6.1          38.0         139.2          18.0
                            2-1-78                     Surface        4.9         30.87         io.9          11.0          66.4
                                                       Bottom         5.0         30.88         10.7          12.0          77.2           6.8
                            4-24-78                    Surface       20.9         33.01          7.8          18.0         125.6          38.4
                                                       Bottom        19.8         33.06          6.9          22.0         154.8          18.0
                            7-24-78                    Surface       30.3         34.38          5.5          17.0         132.8          62.8
                                                       Bottom        29.3         34.35          5.,6         10.0         115.6          51.6

                                                                              10

         Table 1. (Cont.)

                                                                                                                                 SOLIDS
                                      STATION                     TEMP.       SALINITY        D.O.      TURBIDITY
          STATION         DATE         DEPTH         DEPTH         (C)         (0/00)        (mg/1)        (FTU)         TOTAL      SETTLEABLE

           SB12         10-19-77           1         Surface      14.9          34.77          7.2         19.0          109.6          15.2
                                                     Bottom       14.9          34.71          7.4         23.0                -            -
                         2-1-78                      Surface       4.9          30.61         10.6         21.0          124.8          33.2
                                                     Bottom        4.9          30.65         10.4         22.0          163.2          77.2
                         4-25-78                     Surface      21.9          33.14          7.2         17.0          114.8          47.6
                                                     Bottom       21.9          33.15          6.9         64.0          232.0          58.4
                         7-24-78                     Surface      30.0          34.18          5.5         14.0                -            -
                                                     Bottom       29.9          34.18          5.5         20.0          105.6          13.6

           SB13         10-19-77           3         Surface      16.6          34.60          7.2         13.0                -            -
                                                     Bottom       16.5          34.60          7.5         12.0                -            -
                         2-1-78                      Surface       5.0          30.53         10.7         16.0                -            -
                                                     Bottom        4.8          30.57         10.6         21.0                -            -
                         4-24-78                     Surface      21.0          33.04          7.2         24.0                -            -
                                                     Bottom       19.9          33-00          7.1         33.0.         174.8          25.6
                         7-24-78                     Surface      30.0          32.30          5.2         13.0           83.2           0.8
                                                     Bottom       30.0          34.16          5.3         17.0          103.6           6.8

           SB14         10-19-77           2         Surface      17.8          34.35          7.3         14.0           95.6          10.0
                                                     Bottom       17.5          34.35          7.4         19.0          127.6          24.8
                         2-1-78                      Surface       5.3          30.89         10.6           9.7               -            -
                                                     Bottom        5.3          30.93         10.5         10.0           77.2           4.8
                         4-25-78                     Surface      21.3          34.11          6.9         12.0          139.2          23.6
                                                     Bottom       21.0          32.92          7.0         12.0          128.4          21.6
                         7-24-78                     Surface      29.4          34.29          6.0         17.0          122.8          29.2
                                                     Bottom       29.4          34.27          6.1         17.0          126.0          17.2

           SB15          8-22-77           4         Surface      27.8          33.73          7.3         13.0          109.6          31.2
                                                     Bottom       27.0          33.57          5.6         30.0          178.4          34.8
                         2-1-78                      Surface       4.9          30.87         10.4         11.0           95.2          18.0
                                                     Bottom        4.8          30.80         10.6         11.0          100.0          16.8
                         4-24-78                     Surface      20.8          33.01          7.0         11.0           77.6          10.8
                                                     Bottom       19.9          33.00          6.9         33.0          188.4          49.2
                         7-24-78                     Surface      29.7          34.33          5.8         12.0          114.8           8.6
                                                     Bottom       29.3          34.24          6.1         13.0          133.6           7.6

           SB17         10-19-77           1         Surface      16.3          34.71          7.0         18.0          115.6          28.0
                                                     Bottom       16.3          34.72          7.2         17.0          135.6          12.0
                         2-1-78                      Surface       4.9          30.70         10.3         29.0                -            -
                                                     Bottom        4.9          30.73         10.3         32.0                -            -
                         4-25-78                     Surface      21.8          32.79          6.5         16.0          125.2          10.0
                                                     Bottom       21.8          33.13          6.8         20.0          151.6          16.0
                         7-24-78                     Surface      30.1          34.07          5.2         18.0          122.0          21.2
                                                     Bottom       30.1          34.03          5.0         42.0          176.8          24.8

           SBI8         10-19-77           2         Surface      16.8          34.72          7.2         17.0          120.4          29.6
                                                     Bottom       16.7          34.73          7.1         21.0          152.0          22.4
                         2-1-78                      Surface       4.9          30.59         10.6         17.0           78.0          12.4
                                                     Bottom        4.7          30.42         10.6         17.0                -            -
                         4-24-78                     Surface      19.6          33.10          6.8            -          135.6          20.4
                                                     Bottom       19.6          33.06          6.9         22.0          146.8          13.2
                         7-24-78                     Surface      30.1          33.99          5.2         25.0                -            -
                                                     Bottom       30.0          33.97          5.0         51.0          181.2          30.4

          *Samples contained too much silt to make an accurate analysis possible.

                     Table 2. Occurrence of epifaunal invertebrates in dredge collections from station SBO1, Sewee Bay.
                                Collections were taken before, immediately after, and three months after dredging, as well
                                as one year after the pre-dredge survey.

                                Species                      August           February           May          Late July

                     Phylum Porifera
                          Halichondria bowerbanki                                                 +
                          Cliona sp.                           +
                     Phylum Cnidaria
                          Bougainvillia rugosa                                                                     +
                          Pandeidae (undet.)                   +                                  +                +
                          Eudendrium carneum                                                                       +
                          Campanulina sp.                                                         +
                          Clytia kincaidi                                                                          +
                          Obelia bidentata                     +                                  +                +
                          Schizotricha tenella                                                                     +
                          Leptogorgia virgulata                +                                  +                +
                          Actiniaria (undet.)
                     Phylum Entoprocta
                          Barentsia laxa                                                                           +
                     Phylum Bryozoa
                          Alcyonidium hauffi                                                      +
                          Alcyonidium polyoum
                          Anguinella palmata                   +                 +                +                +
                          Amathia distans                                                         +
                          Bowerbankia gracilis                                                    +
                          Membranipora tenuis                  +                 +                +                +
                          Electra monostachys                  +                 +                +
                          Bugula neritina                      +                                  +
                          Schizoporella errata                 +                                  +                +
                          Hippoporina verrilli                 +                                  +                +
                          Parasmittina nitida                                                     +                +
                          Cryptosula pallasi@na                                  +                +                +
                     Phylum Annelida
                          Nereis succinea                                                         +                +
                          Sabellaria vulgaris                  +                 +                +                +
                          Hydroides dianthus                   +
                     Phylum Mollusca
                          Acanthodoris pilosa                                                     +
                          Brachidontes exustus                                                    +
                          Ostrea equestris                     +                                                   +
                     Phylum Artl@ropoda
                          Balanus venustus                     +                                                   +
                          Balanus improvisus                   +                 +
                          Neopanope sayi                                                          +
                          Panopeus herbstii                                                                        +
                     Phylum Echinodermata
                          Sclerodactyla briareus               +
                          Ophiothrix angulata                                                     +                +
                     Phylum Chordata
                          Didemnum. candidum                                                                       +
                          Molgula manhattensis                                   +                                 +

                     No. Species                               16                7                23               23

                     Sorensen Coefficient                                       .43              .51              .51

                                                                         12

                   Table 3. Occurrence of epifaunal invertebrates in dredge collections from station SB03, Sewee Bay.
                             Collections were taken before, immediately after, and three months after dredging, as
                             well as one year after the pre-dredge survey.

                                 Species                      August           February           ApEil           Late July

                   Phylum Porifera
                        Lissodendoryx carolinensis               +                                                      +
                        Microciona prolifera                     +                                  +                   +
                        Halichondria bowerbanki                  +                 +                +                   +
                        Cliona sp.                               +                 +                +                   +
                   Phylum Cnidaria
                        Turritopsis nutricula                    +
                        Bougainvillia rugosa                                                                            +
                        Pandeidae (undet.)                       +                                  +
                        Eudendrium carneum                       +                                  +                   +
                        Campanulina sp.                                                             +
                        Clytia cyli@drica                                                           +
                        Clytia kincaidi                                                                                 +
                        Obelia bidentata                                                            +                   +
                        Obelia dichotoma                                                            +
                        Campano sis sp.                                                             +
                        Dynamen cornicina                        +
                        Sertularia stookeyi                      +                                  +                   +
                        Schizot@@icha tenella                    +                                  +                   +
                        Plumularia floridana                                                                            +
                        Leptogorgia virgulata                    +                 +                +                   +
                        Actiniaria (undet.)                      +                                                      +
                   Phylum Platyhelminthes
                        Stylochus ellipticus                     +
                   Phylum Rhynchocoela
                        Nemertean (undet.)                                         +
                   Phylum Entoprocta
                        Loxosomella sp.                                                             +
                        Pedicellina cernua                                                          +
                        Barentsia laxa                           +                                                      +
                   Phylum Bryozoa
                        Alcyonidium hauffi                       +                 +                4-
                        Anguinella palmata                       +                 +                +                   +
                        Amathia dis_t_.ns                        +                                  +                   +
                        Bowerbankia gracilis                                       +                +                   +
                        Aeverrillia setigera                     +                                  +                   +
                        Membranipora arborescens                                                    +
                        Membranipora tenuis                      +                 +                +                   +
                        Electra monostachys                      +                                  +
                        Synnotum aegyptiacum                     +                                  +                   +
                        Bugula neritina                          +                 +
                        Schizop rella errata                     +                 +                                    +
                        Hippoporina verrilli                     +                 +                +                   +
                        Microporella ciliata                     +                 +                +                   +
                        Parasmittina nitida                      +                 +                +                   +
                   Phylum Annelida
                        Nereis succinea                          +                 +                +
                        Sabellaria vulgaris                      +                 +                +                   +
                        Hydroides dianthus                       +                 +                +                   +
                   Phylum Mollusca
                        Diodora cayenensis                       +
                        Cerithiopsis greeni                                                         +                   +
                        Crepidula fornicata                                                         +                   +
                        Crepidula Plana                                            +
                        Mitrella lunata                                                                                 +
                        Opisthobranchs (undet.)                                                     +
                        Acanthodoris pilosa                      +                 +                +                   +
                        Chaetopleura apiculata                                                      +
                        Ostrea equestris                         +                 +                +                   +
                        Crassostrea virginica                                      +                                    +
                   Phylum Arthropoda
                        Tanystylum orbiculare                    +                                  +
                        Callipallene brevirostrum                                                                       +
                        Balanus  venustus                                          +                +                   +
                        Balanus  galeatus                        +                 +                +
                        Balanus  improvisus                                        +                +                   +

                                                                      13

                         Caprellidae (undet.)                     +                +                +                +
                         Palaemonetes vulgaris                                     +
                         Periclimenes longicaudatus               +
                         Alpheus normanni                         +                +                                 +
                         Clibanarius vittatus                                      +
                         Pagurus longicarpus                                                        +
                         Callinectes ornatus                                                                         +
                         Callinectes sapidus                                       +
                         Menippe mercenaria                       +                                                  +
                         Neopanope sayi                           +                +                +                +
                         Eurypanopeus depressus                                    +                +
                         Panopeus herbstii                                         +                +                +
                         Libinia sp.                              +
                    Phylum Echinodermata
                         Asterias forbesi                                                           +
                         Hemipholis elongata                      +
                         Ophiothrix angulata                      +                +                                 +
                    Phylum Chordata
                         Didemnum candidum                        +                                                  +
                         Perophora viridis                        +                                                  +
                         Molgula manhattensis                     +                                 +                +

                    No. Species                                   45               32               47                46

                    Sorensen Coefficient                                          .55              .61               .68

                                                                       14

                  Table 4. Occurrence of epifaunal invertebrates in dredge collections from station SB04, Sewee Bay.
                            Collections were taken before, immediately after, and three months after dredging, as
                            well as one year after the pre-dredge survey.

                              Species                    August           February          April           Late July

                  Phylum Porifera
                      Microciona prolifera                  +
                      Halichondria bowerbanki               +
                      Cliona sp:                            +                                                   +
                  Phylum Cnidaria
                      Hydractiniidae (undet.)               +
                      Eudendrium carneum                    +                                                   +
                      Clytia kincaidi                                                                           +
                      Dynamena cornicina                                                                        +
                      Sertularia stookeyi                   +
                      Schizotricha tenella                  +                                                   +
                      Plumularia floridana                                                                      +
                      Leptogorgia virgulata                 +                                 +                 +
                      Actiniaria (undet.)                   +
                  Phylum Platyhelminthes
                      Stylochus ellipticus
                  Phylum Entoprocta
                      Loxosomella sp.                                                                           +
                      Barentsia laxa                        +                                                   +
                  Phylum Bryozoa
                      Nolella stipata                       +
                      Anguinella palmata                    +                                                   +
                      Amathia distans                       +
                      Bowerbankia gracilis                  +                                                   +
                      Aeverrillia setigera                                                                      +
                      Membranipora tenuis                   +
                      Synnotum aegyptiacum                  +                                                   +
                      Bugula neritina                       +                                                   +
                      Schizoporella errata                  +                                                   +
                      Hippoporina verrilli                  +                                                   +
                      Microporella ciliata                  +                                                   +
                      Parasmittina nitida                   +                                                   +
                  Phylum Annelida
                      Nereis succinea                                         +
                      Sabellidae (undet.)                   +
                      Hydroides dianthus                                                                        +
                      Oligochaeta (undet.)                                    +
                  Phylum Mollusca
                      Crepidula fornicata                   +                                                   +
                      Simnia uniplicata                     +
                      Anachis avara                         +
                      Busycon canaliculatum                                                   +
                      Ilyanassa obsoleta                                      +
                      Acanthodoris pilosa                   +                                                   +
                      Ostrea equestris                      +                                                   +
                      Crassostrea virginica                 +
                  Phylum Arthropoda
                      Balanus venustus                                                                          +
                      Balanus galeatus                      +                                 +
                      Caprellidae (undet.)                                                    +                 +
                      Alpheus normanni                      +                                                   +
                      Callinectes sapidus                                     +               +
                      Eurypanopeus depressus                +
                      Panopeus herbstii                     +
                      Neopanope sayi                        +
                      Libinia sp.                           +
                  Phylum Echinodermata
                      Sclerodactyla briareus                                  +                                 +
                      Hemipholis elongata                   +                 +
                      Ophiothrix angulata                   +
                  Phylum Chordata
                      Didemnum candidum                     +                                                   +
                      Perophora viridis                                                                         +
                      MoIgula manhattensis                  +                                                   +
                      Ascidiacea (undet.)                   +

                  No. Species                               40                6               5                 29

                  Sorensen Coefficient                                       .04              .09              .55

                                                                  15

                    Table 5. Occurrence of epifaunal invertebrates in dredge collections from station SB05, Sewee Bay.
                               Collections were taken before, immediately after, and three months after dredging, as
                               well as one year after the pre-dredge survey.

                                Species                    Aucnist                                          Late July

                    Phylum Porifera
                         Halichondria bowerbanki                                                +
                         Cliona sp.                           +                                 +
                    Phylum Cnidaria
                         Ectopleura  dumortieri               +
                         Pandeidae (undet.)                                                                     +
                         Eudendrium carneum                   +
                         Clytia kincaidi                      +
                         Obelia bidentata                     +                                 +
                         Sertularia stookeyi                  +
                         Schizotricha tenella                                                   +               +
                         Leptogorgia virguITta                +                                                 +
                         Actiniaria (undet.)                  +                                                 +
                    Phylum Entoprocta
                         Loxosomella sp.                                                        +
                    Phylum Bryozoa
                         Alcyonidium hauffi                   +                                 +
                         Alcyonidium polyoum                  +
                         Anguinella palmata                   +                 +               +               +
                         Amathia distans                      +                                 +
                         Bowerbankia gracilis                 +                                 +               +
                         Aeverrillia setigera                 +                 +
                         Membranipora arborescens             +
                         Membranipora tenuis                  +                                 +               +
                         Electra monostachys                                                    +
                         Bugula neritina                      +                                                 +
                         Schizoporella errata                 +                                 +               +
                         Hippoporina verrilli                 +                                 +
                         Microporella ciliata                                                                   +
                         Parasmittina nitida                  +                                 +               +
                         Cryptosula pallasiana                                                  +               +
                    Phylum Annelida
                         Lepidonotus sublevis                 +
                         Sabellaria vulgaris                  +                                 +               +
                         Hydroides dianthus                   +                                                 +
                    Phylum Mollusca
                         Crepidula fornicata                  +
                         Crepidula plana                      +
                    Phylum Arthropoda
                         Balanus venustus                     +
                         Penaeus aztecus                                                                        +
                         Pagurus longicarpus                                                    +
                         Neopanope sayi                                         +               +
                         Eurypanopeus depressus                                                 +
                    Phylum Echinodermata
                         Sclerodactyla briareus               +
                         Ophiothrix angulata                  +                                 +               +
                    Phylum Chordata
                         Perophora viridis                    +
                         Molgula manhattensis                 +
                         Ascidiacea (undet.)                  +

                    No. Species                               31                3               20               16

                    Sorensen Coefficient                                       .12             .47              .47

                                                                       16

                   Table 6. Occurrence of epifaunal invertebrates in dredge collections from station SB07, Sewee Bay.
                             Collections were taken before, immediately after, and three months after dredging, as well
                             as one year after the pre-dredge survey.

                                Species                       August           February           May           September

                   Phylum Porifera
                        Lissodendoryx carolinensis               +                                 +                +
                        Microciona prolifera                     +                                 +
                        Halichondria bowerbanki                                                    +
                        Cliona sp.                                                 +               +                +
                        Porifera (undet.)                        +
                   Phylum Cnidaria
                        Pandeidae (undet.)                                                         +                +
                        Eudendrium carneum                       +                                 +                +
                        Cuspidella humilis                       +
                        Lovenella grandis                                                          +                +
                        Clytia kincaidi                          +                                 +                +
                        Clytia paulensis                         +                                                  +
                        Obelia bidentata                         +                                                  +
                        Campanopsis sp.                                                            +
                        Dynamena cornicina                       +
                        Sertularia stookeyi                      +
                        Schizotricha tenella                                                       +                +
                        Leptogorgia virgulata                    +                 +               +                +
                        Actiniaria (undet.)                      +                 +               +                +
                   Phylum Entoprocta
                        Loxosomella sp.                                                            +
                        Barentsia laxa                           +                                 +                +
                   Phylum Bryozoa
                        Alcyonidium hauffi                                         +               +
                        Anguinella palmata                       +                 +               +                +
                        Amathia distans                          +                 +               +
                        Bowerbankia gracilis                                       +               +                +
                        Aeverrillia setigera                     +                 +                                +
                        Membranipora arborescens                                                   +
                        Membranipora tenuis                      +                 +                                +
                        Electra monostachys                                        +               +
                        Synnotum aegyptiacum                     +                                 +
                        Bugula neritina                          +                 +               +
                        Schizoporella errata                     +                 +               +                +
                        Hippoporina verrilli                     +                 +               +                +
                        Microporella ciliata                                       +               +                +
                        Parasmittina nitida                      +                 +               +                +
                   Phylum Annelida
                        Nereis succinea                                            +                                +
                        Sabellidae (undet.)                                        +
                        Sabellaria vulgaris                      +                 +               +                +
                        Hydroides dianthus                       +                 +               +                +
                   Phylum Mollusca
                        Crepidula fornicata                                                        +
                        Eupleura caudata                                                           +
                        Cerithiopsis greeni                                                        +
                        Ilyanassa obsoleta                                                                          +
                        Acanthodoris pilosa                                                        +                +
                        Chaetopleura apiculata                                                     +
                        Brachidontes exustus                                       +
                        Ostrea equestris                         +                 +               +
                        Crassostrea virginica                                      +                                +
                   Phylum Arthropoda
                        Callipallene brevirostrum                                                  +
                        Balanus galeatus                                                                            +
                        Balanus improvisus                                         +               +                +
                        Anthuridae (undet.)                                        +
                        Caprellidae (undet.)                                       +               +
                        Penaeus duorarum                         +
                        Alpheus armillatus                                         +
                        Alpheus normanni                                                           +                +
                        Pagurus longicarpus                                                        +
                        Callinectes sapidus                                                        +
                        Hexapanopeus angustifrons                                  +

                                                               17

                      Table 6. (cont.)

                                   Species                       August          February           May           September

                           Neopanope sayi                                            +               +                +
                           Eurypanopeus depressus                                                    +
                           Panopeus herbstii                        +                                +
                           Libinia sp.                                               +               +
                      Phylum Echinodermata
                           Ophiothrix angulata                      +                +               +                +
                      Phylum Chordata
                           Didemnum candidum                                                         +                +
                           Perophora viridis                        +                                +                +
                           Molgula manhattensis                     +                +               +                +

                      No. Species                                   30               31              48               34

                      Sorensen Coefficient                                           .49            .54              .59

                                                                       18

                   Table 7. Occurrence of epifaunal invertebrates in dredge collections from station SB08, Sewee Bay.
                             Collections were taken before, immediately after, and three months after dredging, as
                             well as one year after the pre-dredge survey.

                               Species                        August           February            May          Late July

                   Phylum Porifera
                        Lissodendoryx carolinensis                +                                                   +
                        Microciona prolifera                      +                 +                +                +
                        Halichondria bowerbanki                                     +                +                +
                        Cliona sp.                                +                 +                +                +
                   Phylum Cnidaria
                        Turritopsis nutricula                                                        +
                        Bougainvillia rugosa                                                                          +
                        Pandeidae (undet.)                                                           +
                        "Perigonimus' sp.                         +
                        Eudendrium carneum                        +                                  +                +
                        Lovenella grandis                                           +
                        Clytia kincaidi                           +                                  +                +
                        Obelia bidentata                          +                                                   +
                        Schizotricha tenella                      +                                  +                +
                        Plumularia floridana                                                                          +
                        Leptogorgia virgulata                     +                                  +                +
                        Actiniaria (undet.)                                                                           +
                   Phylum Entoprocta
                        Loxosomella sp.                                                                               +
                        Pedicellina cernua                                                           +
                        Barentsia laxa                            +                                  +                +
                        Barentsia sp.                                                                +
                   Phylum Bryozoa
                        Crisia sp.                                                                   +
                        Anguinella   palmata                      +                 +                +
                        Bowerbankia gracilis                      +                 +                +                +
                        Aeverrillia setigera                      +                                                   +
                        Membranipora arborescens                                                                      +
                        Membranipora tenuis                       +                 +                +                +
                        Conopeum tenuissimum                                                                          +
                        Electra monostachys                                         +
                        Synnotum aegyptiacum                      +
                        Bugula neritina                           +                                  +                +
                        Schizoporella errata                      +                 +                +                +
                        Hippoporina verrilli                      +                 +                +                +
                        Microporella ciliata                      +                 +                +                +
                        Parasmittina nitida                       +                 +                +                +
                   Phylum Annelida
                        Nereis succinea                                             +                +
                        Lepidonotus sublevis                                                         +
                        Sabellaria @@@                            +                 +                +
                        Hydroides dianthus                        +                                  +                +
                        Polydora sp.                                                +
                   Phylum Mollusca
                        Crepidula fornicata                       +                 +                                 +
                        Crepidula Plana                                             +
                        Simnia uniplicata                                                            +
                        Urosalpinx cinerea                                                           +
                        Acanthodoris pilosa                                                                           +
                        Ostrea equestris                          +                 +                                 +
                        Crassostrea virginica                     +                 +                +                +
                   Phylum Arthropoda
                        Tanystylum orbiculare                                                                         +
                        Balanus venustus                          +                                  +
                        Balanus improvisus                                          +                +                +
                        Caprellidae (undet.)                                                         +                +
                        Penaeus aztecus                                                                               +
                        Penaeus uorarum                           +
                        Portunus gibbesii                         +
                        Callinectes ornatus                       +
                        Callinectes sapidus                       +
                        Menippe mercenaria                        +                                  +
                        Hexapanopeus angustifrons                                   +
                        Neopanope sayi                                                               +
                        Eurypanopeus depressus                    +                                  +                +
                        Panopeus herbstii                                           +                +                +

                                                                    19

                     Table 7. (cont.)

                                 Species                      August          February          May           Late July

                     Phylum Echinodermata
                          Sclerodactyla briareus                                                 +
                          Ophiothrix angulata                     +                +             +
                     Phylum Chordata
                          Perophora viridis                       +                              +                +
                          Molgula manhattensis                    +                +             +

                     No. Species                                 35               24             40              36

                     Sorensen Coefficient                                        .51             .67            .65

                                                                     20

                  Table 8. Occurrence of epifaunal invertebrates in dredge collections from station SB10, Sewee Bay,
                             Collections were taken before, immediately after, and three months after dredging, as
                            well as one year after the pre-dredge survey.

                              Species                       August          Feburary           April           Late July

                  Phylum Porifera
                       Microciona prolifera                                                                        +
                       Halichondria bowerbanki                 +                                                   +
                       Cliona sp.                              +
                  Phylum Cnidaria
                       Turritopsis nutricula                                                                       +
                       Pandeidae (undet.)                      +
                       Eudendrium carneum                                                                          +
                       Clytia kincaidi                         +                                                   +
                       Obelia bidentata                        +
                       Dynamena cornicina                      +                                                   +
                       Sertularia stookeyi                     +
                       Schizotricha tenella                                                       +
                       Leptogorgia virgulata                   +                                                   +
                       Actiniaria (undet.)                     +
                  Phylum Entoprocta
                       Barentsia laxa                          +                                                   +
                  Phylum Bryozoa
                       Anguinella palmata                      +                                                   +
                       Amathia distans                         +
                       Bowerbankia gracilis                    +                                  +                +
                       Aeverrillia setigera                    +                                                   +
                       Membranipora arborescens                +
                       Membranipora tenuis                     +                 +                                 +
                       Electra monostachys                                                        +
                       Bugula neritina                         +                                                   +
                       Schizoporella errata                    +                                                   +
                       Hippoporina verrilli                    +                                                   +
                       Microporella ciliata                    +                                                   +
                       Parasmittina nitida                     +
                       Cryptosula pallasiana                   +
                  Phylum Annelida
                       Sabellaria vulgaris                     +                                                   +
                       Hydroides dianthus                      +                                                   +
                  Phylum Mollusca
                       Simnia uniplicata                       +
                       Ilyanassa obsoleta                                        +
                       Ostrea equestris                                                                            +
                  Phylum Arthropoda
                       Balanus venustus                        +                                                   +
                       Balanus improvisus                                                                          +
                       Pagurus longicarpus                                       +
                       Eurypanopeus depressus                  +                                  +                +
                  Phylum Echinodermata
                       Hemipholis elongata                                                                         +
                       Ophiothrix angulata                                                                         +
                  Phylum Chordata
                       Perophora viridis                                                                           +
                       Molgula manhattensis                    +                                                   +

                  No. Species                                  28                3                4                26

                  Sorensen Coefficient                                          .06               13              .67

                                                                      21

                          Table 9. Occurrence of epifaunal invertebrates in dredge collections from station SB12,
                                    Sewee Bay. Collections were taken before, immediately after and three months
                                    after dredging, as well as one year after the pre-dredge survey.

                                   Species                    August           February           May           September

                          Phylum Mollusca
                               Nassarius vibex                   +
                               Ilyanassa obsoleta                                  +                                 +
                          Phylum Arthropoda
                               Penaeus aztecus                                                     +
                               Callinectes sapidus               +                 +
                          Phylum Chordata
                               Molgula manhattensis              +

                          No. Species                            3                 2               1                 1

                          Sorensen Coefficient                                  0.40            0.00              0.00

                                                                        22

                    Table 10. Occurrence of epifaunal invertebrates in dredge collections from station SB13,
                                Sewee Bay. Collections were taken before, immediately after, and three months
                                after dredging, as well as one year after the pre-dredge survey.

                                  Species                        August            February            April           Late July

                    Phylum Porifera
                          Lissodendoryx carolinensis                 +
                          Microciona prolifera                       +
                          Cliona sp.                                 +                 +                 +                 +
                    Phylum Cnid aria
                          Ectopleura  dumortieri                                       +
                          Hydractiniidae (undet.)                    +                                   +
                          Pandeidae (undet.)                         +                 +                 +                 +
                          Amphinema dinema                                                               +
                          Eudendrium carneum                         +                                   +                 +
                          Campanulina sp.                                                                                  +
                          Lovenella grandis                                                                                +
                          Clytia kincaidi                            +                                   +                 +
                          Clytia paulensis                                                               +
                          Obelia bidentata                                                                                 +
                          Obelia dichotoma                                                                                 +
                          Dynamena cornicina                                                                               +
                          Schizotricha tenella                       +                                   +                 +
                          Leptogorgia virgulata                      +                 +                                   +
                          Actiniaria (undet.)                        +                                                     +
                    Phylum Entoprocta
                          Pedicellina cernua                                           +                 +
                          Barentsia laxa                             +                                   +                 +
                    Phylum Bryozoa
                          Alcyonidium hauffi                                                                               +
                          Anguinella palmata                         +                 +                 +                 +
                          Bowerbankia gracilis                       +                 +                 +                 +
                          AeverrTllia setigera                       +                                                     +
                          Membranipora arborescens                                                       +
                          Membranipora tenuis                        +                 +                 +                 +
                          Electra monostachys                                                            +
                          Synnotum aegyptiacum                       +
                          Schizoporella errata                       +                 +                 +                 +
                          Hippoporina verrilli                       +                 +                 +                 +
                          Microporella ciliata                       +                 +                 +                 +
                          Parasmittina nitida                        +                                                     +
                    Phylum Annelida
                          Nereis succinea                                              +                 +
                          Lepidonotus sublevis                                                                             +
                          Sabellaria vulgaris
                          Hydroides dianthus                         +                                   +                 +
                          Sabellidae (undet.)                        +                 +
                    Phylum Mollusca
                          Littorina irrorata                                                                               +
                          Crepidula fornicata                                          +
                          Crepidula plana                                                                                  +
                          Simnia uniplicata                                                                                +
                          Urosalpinx cinerea                         +
                          Ilyanassa obsoleta                                           +                 +
                          Acanthodoris pilosa                        +                                                     +
                          Ostrea equestris                           +                 +                                   +
                          Crassostrea virginica                      +
                    Phylum Arthropoda
                          Callipallene brevirostrum                  +
                          Balanus galeatus                                                                                 +
                          Balanus improvisus                                                             +                 +
                          Cyathura burbancki                                           +
                          Penaeus aztecus                                                                                  +
                          Clibanarius vittat-us.                                       +
                          Callinectes sapidus                                          +
                          Menippe mercenaria                         +
                          Hexapanopeus angustifrons                                    +
                          Panopeus herbstii                          +
                    Phylum Echinodermata
                          Sclerodactyla briareus                                       +
                          Ophiothrix angulata                        +                                                     +

                                                                        23

                   Table 10. (cont.)

                                Species                      August          February          April          Late July

                   Phylum Chordata
                        Didemnum candidum                                                                          +
                        Perophora viridis                                                                          +
                        Molgula manhattensis                    +                +               +                 +

                   No. Species                                 31               22              25                37

                   Sorensen Coefficient                                        .45             .57                .62

                                                                    24

                   Table 11. Occurrence of epifaunal invertebrates in dredge collections from station SB14, Sewee
                               Bay. Collections were taken before, immediately after, and three months after dredging,
                               as well as one year after the pre-dredge survey.

                                Species                    August            February           April           Late July

                   Phylum Porifera
                         Microciona prolifera                  +                 +                +                 +
                         Cliona sp.                            +                 +                +                 +
                   Phylum Cnidaria
                         Ectopleura   dumortieri                                                                    +
                         Bougainvillia rugosa                                                                       +
                         Pandeidae (undet.)                                                       +
                         tudendrium carneum                                                       +                 +
                         Campanulina sp.                                         +                +                 +
                         Lovenella gracilis                                                       +                 +
                         Lovenella grandis                                       +                +                 +
                         Clytia kincaidi                       +                                                    +
                         Obelia bidentata                      +                                                    +
                         Obelia dichotoma                                                                           +
                         @@@ SP.                                                                                    +
                         Schizotricha tenella                  +                                                    +
                         Leptogorgia virgulata                 +                 +                +                 +
                         Actinia:ria (undet.)                                    +                +                 +
                   Phylum Platyhelminthes
                         Stylochus ellipticus                  +                 +
                   Phylum Rhynchocoela
                         Nemertean (undet.)                                      +
                   Phylum Entoprocta
                         Barentsia laxa                        +                 +                +                 +
                   Phylum Bryozoa
                         Alcyonidium hauffi                                                       +
                         Alcyonidium mammillatum                                 +
                         Anguinella palmata                    +                 +                +                 +
                         Arnathia distans                      +                 +
                         Bowerbankia gracilis                  +                 +                +                 +
                         Aeverrillia setigera                  +                 +                +                 +
                         Membranipora arborescens              +
                         Membranipora tenuis                   +                 +                +                 +
                         Electra monostachys                                     +                +
                         Bugula neritina                       +                                  +                 +
                         Schizoporella errata                  +                 +                +                 +
                         Hippoporina verrilli                  +                 +                +                 +
                         Parasmittina nitida                   +                 +                +                 +
                         Cryptosula pallasiana                                                    +                 +
                   Phylum Annelida
                         Nereis succinea                                         +
                         Sabellaria vulgaris                   +                 +                +                 +
                         Hydroides dianthus                    +                 +                +                 +
                         Polydora sp.                                            +
                   Phylum Mollusca
                         Crepidula fornicata                                     +                                  +
                         Simnia uniplicata                     +
                         Acanthodoris pilosa                                     +                +                 +
                         Brachidontes exustus                                    +                +
                         Ostrea equestris                      +                 +                +                 +
                         Crassostrea virginica                 +                 +
                   Phylum Arthropoda
                         Tanystylum orbiculare                 +
                         Balanus venustus                      +                 +
                         Balanus galeatus                      +
                         Balanus improvisus                    +                 +                +                 +
                         Caprellidae (undet.)                                    +                +
                         Penaeus aztecus                       +                                                    +
                         Palaemonetes vulgaris                                                    +
                         Clibanarius vittatus                  +
                         Callinectes sapidus                                     +                +

                                                                        25

                      Table 11. (cont.)

                                   Species                 August           February          April          Late July

                           Neopanope sayi                                                       +
                           Eurypanopeus depressus                               +               +
                           Panopeus herbstii                                    +               +
                      Phylum Echinodermata
                           Ophiothrix angulata                                                  +
                      Phylum Chordata
                           Didemnum candidum                  +                 +               +
                           Perophora viridis                                                    +                 +
                           Molgula manhattensis               +                 +               +                 +

                      No. Species                             31                36              38               34

                      Sorensen Coefficient                                     .63             .55              .65

                                                                     26

                      Table 12. Occurrence of epifaunal invertebrates in dredge collections from station SB15,
                                  Sewee Bay. Collections were taken before, immediately after, and three months
                                  after dredging, as well as one year after the pre-dredge survey.

                                  Species                     August           February           May           Late July

                      Phylum Porifera
                           Microciona prolifera                   +
                           Halichondria bowerbanki                +                                +                +
                           Cliona sp.                             +                                +                +
                      Phylum Cnidaria
                           Turritopsis nutricula                                                                    +
                           Pandeidae (undet.)                                                                       +
                           Eudendrium album                                                        +
                           Campanulina sp.                                         +                                +
                           Clytia kincaidi                                                         +                +
                           Obelia bidentata                       +                                                 +
                           Dynamena cornicina                     +
                           Sertularia stookeyi                    +                                +
                           Schizotricha tenella                   +                                +                +
                           Leptogorgia virgulata                  +                                +                +
                           Astrangia danae                        +
                           Actiniaria (undet.)                    +                                +                +
                      Phylum Entoprocta
                           Barentsia laxa                         +                                +                +
                      Phylum Bryozoa
                           Alcyonidium hauffi                     +                                +                +
                           Alcyonidium polyoum                                                                      +
                           Anguinella palmata                     +                +               +                +
                           Bowerbankia gracilis                   +                +
                           Aeverrillia setigera                   +                                +                +
                           Membranipora arborescens               +
                           Membranipora tenuis                    +                +               +                +
                           Electra monostachys                                     +               +
                           Bugula neritina                        +                                +                +
                           Schizoporella errata                   +                                +                +
                           Hippoporina verrilli                   +                                +
                           Microporella ciliata                   +                                +
                           Parasmittina nitida                    +                +               +
                           Cryptosula pallasiana                                                   +
                      Phylum Annelida
                           Nereis succinea                                                         +
                           Lepidonotus sublevis                                                    +
                           Sabellaria vulgaris                    +                +
                           Hydroides dianthus                     +
                      Phylum Mollusca
                           Simnia uniplicata                      +
                           Anadara transversa                     +
                           Brachidontes exustus                   +
                           Pteria colymbus                        +
                           Ostrea equestris                       +                                                 +
                      Phylum Arthropoda
                           Balanus venustus                       +
                           Balanus galeatus                                                        +
                           Balanus improvisus                     +                +               +                +
                           Penaeus aztecus                        +
                           Penaeus duorarum                                        +
                           Alpheus normanni                                                        +
                           Pagurus longicarpus                    +
                           Neopanope sayi                                                          +
                           Eurypanopeus depressus                                                  +                +
                           Squilla empusa                                                                           +
                      Phylum Echinodermata
                           Hemipholis elongata                                     +
                           Ophiothrix angulata                    +                                +                +
                      Phylum Chordata
                           Didemnum candidum                                                       +
                           Perophora viridis                                                                        +
                           Molgula manhattensis                   +                +               +
                           Ascidiacea (undet.)                    +                                +

                      No. Species                                 36               11              31               24

                      Sorensen Coefficient                                         .30            .60               .53

                                                                     27

                       Table 13. Occurrence of epifaunal invertebrates in dredge collections from station SB17,
                                   Sewee Bay. Collections were taken before, immediately after, and three months
                                   after dredging, as well as one year after the pre-dredge survey.

                                   Species                    August           February           May           September

                       Phylum Porifera
                             Microciona prolifera                                                  +
                       Phylum Cnidaria
                             Pandeidae (undet.)                  +
                             Eudendrium carneum                  +                                                   +
                             Lovenella grandis                   +
                             Clytia kincaidi                     +
                             Schizotricha tenella                +                                                   +
                             Leptogorgia virgulata               +                                                   +
                       Phylum Bryozoa
                             Victorellidae (undet.)                                                                  +
                             Anguinella palmata                  +                  +                                +
                             Amathia distans                                        +              +
                             Bowerbankia gracilis                +                                 +                 +
                             Aeverrillia setia@@                                                                     +
                             Membranipora tenuis                 +
                             Parasmittina nitida                 +
                       Phylum Molluscs
                             Ilyanassa obsoleta                                     +              +
                       Phylum Arthropoda
                             Callinectes sapidus                 +                  +
                             Panopeus herbstii                                                     +
                       Phylum Echinodermata
                             Ophiothrix angulata                                                   +
                       Phylum Chordata
                             Perophora viridis                                                                       +
                             MoIgula manhattensis                +                  +

                       No. Species                                12                5              6                 8

                       Sorensen Coefficient                                      0.35            0.11             0.50

                                                                         28

                    Table 14. Occurrence of epifaunal invertebrates in dredge collections from station SB18, Sewee
                               Bay. Collections were taken before, immediately after, and three months after
                               dredging, as well as one year after the pre-dredge survey.

                                Species                          August            February            April           Late July

                    Phylum Porifera
                         Lissodendoryx carolinensis                   +                  +
                         Microciona prolifera                         +                                   +
                         Halichondria bowerbanki                      +                  +                                   +
                         Cliona sp.                                   +                  +                +                  +
                         Porifera (undet.)                                               +                                   +
                    Phylum Cnidaria
                         Ectopleura dumortieri                                           +
                         Turritopsis nut.ricula                       +
                         Pandeid-ae- (i  et.)                                                             +                  +
                         Eudendrium carneum                           +                                                      +
                         Campanulina sp.                              +                                   +
                         Lovenella gracilis                                                               +
                         Lovenella grandis                                                                +                  +
                         Clytia cylindrica                                               +
                         Clytia kincaidi                              +                                   +                  +
                         Obelia bidentata                                                                                    +
                         Schizotricha tenella                         +                                   +                  +
                         Plumularia floridana                         +
                         Leptogorgia virgulata
                         Actiniaria (undet.)                          +                  +                +                  +
                    Phylum Entoprocta
                         Pedicellina cernua                           +                                   +
                         Barentsia laxa                               +                                   +                  +
                    Phylum Bryozoa
                         Alcyonidium hauffi                                              +
                         Anguinella palmata                           +                  +                +                  +
                         Amathia distans                              +                  +                +                  +
                         Bowerbankia gracilis                         +                  +                +                  +
                         Aeverrillia setigera                         +
                         Membranipora arborescens                                                         +
                         Membranipora tenuis                          +                  +                +                  +
                         Electra monostachys                                             +                +
                         Synnotum aegyptiacum                         +                  +                                   +
                         Bugula neritina                                                                                     +
                         Schizoporella errata                         +                  +                +                  +
                         Hippoporina verrilli                         +                  +                +                  +
                         Microporella ciliata                         +                  +                +                  +
                         Parasmittina nitida                          +                  +                +                  +
                    Phylum Annelida
                         Nereis succinea                                                 +
                         Lepidonotus sublevis                                            +                +
                         Sabellaria vulgaris                                             +                +                  +
                         Hydroides dianthus                           +                  +                +                  +
                    Phylum Mollusca
                         Crepidula fornicata                          +                                   +
                         Simnia uniplicata                                                                +
                         Eupleura caudata                                                +
                         Fasciolaria lilium hunteria                  +                                                      +
                         Nassarius vibex                                                                  +
                         Acanthodoris pilosa                          +                  +                +
                         Anadara transversa                                              +
                         Ostrea equestris                             +                                                      +
                    Phylum Arthropoda
                         Tanystylum orbiculare                        +                                                      +
                         Balanus galeatus                             +                                   +
                         Balanus improvisus                                              +                +                  +
                         Caprellidae (undet.)                         +                                   +
                         Penaeus duorarum                             +
                         Trachypenaeus constrictus                                                        +
                         Alpheus heterochaelis                        +
                         Alpheus normanni                                                                                    +
                         Portunus gibbesii                            +
                         Callinectes sapidus                                             +                +
                         Neopanope sayi                                                  +
                         Eurypanopeus depressus                                                                              +
                         Panopeus herbstii                                                                                   +

                                                                         29

                   Table 14. (cont.)

                             Species                         August          February          April           Late July

                   Phylum Echinodermata
                       Ophiothrix angulata                      +                 +               +
                   Phylum Chordata
                       Didemnum.candidum                                                                            +
                       Perophora viridis                                                                            +
                       Molgula manhattensis                     +                 +               +                 +
                       Ascidiacea (undet.)                                        +

                   No. Species-                                37                32              36                34

                   Sorensen Coefficient                                         .52              .66              .62

                                                                     30

                  Table 15. Macroinvertebrate species collected from the Sewee Bay channel station, SBOl. (A = Amphipoda;
                             C = Cumacea; D - Decapoda; H = Hemichordata; I = Isopoda; M = Mollusca; My = Mysidacea;
                             0 = Ophiuroidea; P = Polychaeta).

                                                                        SBOl
                                                                                                                            Rank
                                                            Total           Number/0.05 m2            Estimated              by
                           Species                         Number             Y          SD           Number/m2            Number

                  -----------------------------------   Collection Date     23 August 1977  ------------------------------------

                  Paraprionospio pi-anata (P)                 38              12.7       2.3               230               1.0
                  Ogyrides limicola (D)                       10              3.3        1.2               61                2.0
                  Lumbrineris tenuis (P)                        6             2.0        1.0               36                3.0
                  Notomastus lobatus (P)                        4             1.3        1.5               24                4.0
                  Glycinde solitaria (P)                        3             1.0        1.0               18                5.0
                  Aricidea fragilis M                           2             0.7        0.6               12                6.0
                  Saccoglossus kowalevskii (H)                  1             0.3        0.6                 6               8.0
                  Glycera americana (P)                         1             0.3        0.6                 6               8.0
                  Haploscoloplos fragilis (P)                   1             0.3        0.6                 6               8.0

                  -----------------------------------   Collection Date     16 January 1978 -----------------------------------

                  Paraprionospio pinnata (P)                  14              4.7        2.5               85                1.0
                  Ogyrides limicola (D)                       11              3.7        2.3               67                2.0
                  Gattyana cirrosa (P)                          2             0.7        1.2               12                3.5
                  Aricidea fragilis M                           2             0.7        1.2               12                3.5
                  Mysidopsis bigelowi (My)                      1             0.3        0.6                 6               6.0
                  Amphiodia planispina (0)                      1             0.3        0.6                 6               6.0
                  Glycera americana (P)                         1             0.3        0.6                 6               6.0

                  -----------------------------------   Collection Date     28 April 1978  ------------------------------------

                  Melita appendiculata.(A)                    26              8.7       15.0               158               1.0
                  Paraprionospio pinnata (P)                  15              5.0        6.0               91                2.0
                  Nereis succinea (P)                           4             1.3        2.3               24                3.0
                  Eurypanopeus depressus (D)                    2             0.7        1.2               12                7.5
                  Paracaprella tenuis (A)                       2             0.7        1.2               12                7.5
                  Ophiothrix angulata (0)                       2             0.7        1.2               12                7.5
                  Mulinia lateralis (M)                         2             0.7        1.2               12                7.5
                  Scoloplos sp. (P)                             2             0.7        0.6               12                7.5
                  Lumbrineris tenuis M                          2             0.7        1.2               12                7.5
                  Haploscoloplos fragilis (P)                   2             0.7        1.2               12                7.5
                  Sabella microphthalma   (P)                   2             0.7        1.2               12                7.5
                  Ogyrides limicola (D)                         1             0.3        0.6                 6              19.5
                  Neopanope texana (D)                          1             0.3        0.6                 6              19.5
                  Ampelisca vadorum (A)                         1             0.3        0.6                 6              19.5
                  Lembos websteri (A)                           1             0.3        0.6                 6              19.5
                  Leucon americanus (C)                         1             0.3        0.6                 6              19.5
                  Cyathura burbanki (1)                         1             0.3        0.6                 6              19.5
                  Actiniaria (undet.)                           1             0.3        0.6                 6              19.5
                  Glycera americans (P)                         1             0.3        0.6                 6              19.5
                  Heteromastus filiformis (P)                   1             0.3        0.6                 6              19.5
                  Glycinde solitaria (P)                        1             0.3        0.6                 6              19.5
                  Marphysa sanguinea (P)                        1             0.3        0.6                 6              19.5
                  Diopatra cuprea (F)                           1             0.3        0.6                 6              19.5
                  Arabella iricolor (P)                         1             0.3        0.6                 6              19.5
                  Drilonereis magna (P)                         1             0.3        0.6                 6              19.5
                  Aricidea fragilis (P)                         1             0.3        0.6                 6              19.5
                  Cirratulidae (undet.) M                       1             0.3        0.6                 6              19.5

                  ------------------------------------   Collection Date     3 August 1978  -----------------------------------

                  Ogyrides limicola (D)                       21              7.0        4.4               127               1.0
                  Heteromastus filiformis M                     2             0.7        0.6               12                3.0
                  Lumbrineris t-en-uis(P)                       2             0.7        0.6               12                3.0
                  Haploscoloplos fragilis (P)                   2             0.7        1.2               12                3.0
                  Mytilidae(undet.)(M)                          1             0.3        0.6                 6               6.0
                  Nereis succinea (P)                           1             0.3        0.6                 6               6.0
                  Paraprionospio pinnata M                      1             0.3        0.6                 6               6.0

                                                                        31

                     Table 16. Macroinvertebrate species collected from the Sewee Bay intertidal station, SB02. (A
                                Amphipoda; D = Decapoda; I = Isopoda; M = Molluscs; P = Polychaeta).

                                                                           SB02
                                                                                             2                                Rank
                                                              Total            Number/0.05m            Estimated               by
                                 Species                     Number             7         SD           Number/m2             Number

                     -----------------------------------   Collection Date     30 August  1977 ----------------------------------

                     Lumbrineris tenuis (P)                      17             5.7       3.5               103                1.0
                     Aricidea fragilis (P)                       15             5.0       2.0               91                 2.0
                     Glycera americana (P)                        3             1.0       1.0               18                 3.0
                     Alpheus normanni (D)                         2             0.7       1.2               12                 4.0
                     Scoloplos sp. (P)                            1             0.3       0.6                 6                6.0
                     Glycinde solitaria (P)                       1             0.3       0.6                 6                6.0
                     Haploscoloplos fragilis (P)                  1             0.3       0.6                 6                6.0

                     -----------------------------------   Collection Date     16 January 1978-7  -------------------------------

                     Heteromastus filiformis     (P)              8             2.7       2.1               48                 1.0
                     Lumbrineris tenuis (P)                       7             2.3       2.1               42                 2.0
                     Mercenaria mercenaria (M)                    5             1.7       0.6               30                 3.0
                     Glycera americans (P)                        4             1.3       0.6               24                 4.0
                     Tellinidae (undet.) (M)                      2             0.7       0.6               12                 5.0
                     Nemertina (undet.)                           1             0.3       0.6                 6                7.5
                     Turbonilla sp. (M)                           1             0.3       0.6                 6                7.5
                     Paraprionospio pinnata (P)                   1             0.3       0.6                 6                7.5
                     Maldanidae (undet.) (P)                      1             0.3       0.6                 6                7.5

                     ------------------------------------   Collection Date =   25 April  1978 ---------------------------------

                     Nereis succinea (P)                         88            29.3      19.3               533                1.0
                     Uca sp. (D)                                  1             0.3       0.6                 6                2.5
                     Cyathura burbanki (1)                        1             0.3       0.6                 6                2.5

                     ------------------------------------   Collection Date =   3 August  1978 ---------------------------------

                     Scoloplos rubra (P)                          3             1.0       1.0               18                 1.5
                     Heteromastus filiformis (P)                  3             1.0       1.7               18                 1.5
                     Alpheus heterochaelis   '(D)                 2             0.7       1.2               12                 4.0
                     Lumbrineris tenuis (P)                       2             0.7       1.2               12                 4.0
                     Haploscoloplos fragilis (P)                  2             0.7       1.2               12                 4.0
                     Gammarus palustris (A)                       1             0.3       0.6                 6                7.5
                     Glycera americana (P)                        1             0.3       0.6                 6                7.5
                     Scoloplos sp. (P)                            1             0.3       0.6                 6                7.5
                     Drilonereis magna    (P)                     1             0.3       0.6                 6                7.5

                                                                            32

                  Table 17. Macroinvertebrate species collected from the Sewee Bay channel station, SB03. (A
                            Amphipoda; C = Cumacea; D = Decapoda; I = Isopoda; M = Mollusca; P = Polychaeta).

                                                                        SB03
                                                                                                                          Rank
                                                          Total             Number/0.05m 2         Estimated              by
                              Species                     Number            -9        SD           Number/m2              Number

                  ------------------------------------  Collection Date = 30 August 1977   ------------------------------------

                  Lumbrineris tenuis (P)                    26              8.7       3.1             158                 1.0
                  Diptera (undet.)                            1             0.3       0.6                6                2.5
                  Glycera americana (P)                       1             0.3       0.6                6                2.5

                  ------------------------------------  Collection Date     16 January 1978  -----------------------------------

                  Heteromastus filiformis (P)                 4             1.3       0.6              24                 1.5
                  Lumbrineris tenuis (P)                      4             1.3       1.2              24                 1.5
                  Caprella equilibra (A)                      3             1.0       1.7              18                 3.5
                  Paraprionospio pinnata   (P)                3             1.0       0.0              18                 3.5
                  Melita dentata (A)                          2             0.7       1.2              12                 5.5
                  Nereis succinea (P)                         2             0.7       0.6              12                 5.5
                  Alpheus heterochaelis   '(D)                1             0.3       0.6                6                11.0
                  Pinnixa sayana (D)                          1             0.3       0.6                6                11.0
                  Cyathura burbanki (1)                       1             0.3       0.6                6                11.0
                  Mitrella lunata (M)                         1             0.3       0.6                6                11.0
                  Glycera americana (P)                       1             0.3       0.6                6                11.0
                  Glycinde solitaria (P)                      1             0.3       0.6                6                11.0
                  Arabella iricolor (P)                       1             0.3       0.6                6                11.0
                  Aricidea fragilis (P)                       1             0.3       0.6                6                11.0
                  Cirratulidae (undet.)   (P)                 1             0.3       0.6                6                11.0

                  -------------------------------------   Collection Date   = 24 April 1978 ------------------------------------

                  Ampelisca vadorum (A)                     18              6.0       1.0              109                1.0
                  Lumbrineris tenuis (P)                    16              5.3       2.3              97                 2.0
                  Streblospio benedicti   (P)                 5             1.7       2.9              30                 3.0
                  Mulinia lateralis (A)                       4             1.3       0.6              24                 5.0
                  Tellinidae (undet.)(M)                      4             1.3       1.5              24                 5.0
                  Mediomastus californiensis (P)              4             1.3       2.3              24                 5.0
                  Erichthonius brasiliensis (A)               3             1.0       1.7              18                 8.5
                  Glycera americana (P)                       3             1.0       1.7              18                 8.5
                  Tharyx marioni (P)                          3             1.0       1.0              18                 8.5
                  Haplosco,loplos sp. (P)                     3             1.0       1.0              18                 8.5
                  Ogyrides limicola (D)                       2             0.7       0.6              12                 13.5
                  Oxyurostylis smithi (C)                     2             0.7       0.6              12                 13.5
                  Nemertina (undet.)                          2             0.7       0.6              12                 13.5
                  Pelecypoda (undet.) (M)                     2             0.7       0.6              12                 13.5
                  Glycinde solitaria (P)                      2             0.7       0.6              12                 13.5
                  Paraprionospio pinnata   (P)                2             0.7       0.6              12                 13.5
                  Portunidae (undet.) (D)                     1             0.3       0.6                6                20.0
                  Caprellidae (undet.) (A)                    1             0.3       0.6                6                20.0
                  Acteocina canaliculata (M)                  1             0.3       0.6                6                20.0
                  Heteromastus filiformis (P)                 1             0.3       0.6                6                20.0
                  Nereis succinea (P)                         1             0.3       0.6                6                20.0
                  Aricidea fragilis (P)                       1             0.3       0.6                6                20.0
                  Maldanidae (undet.) (P)                     1             0.3       0.6                6                20.0

                  ------------------------------------  Collection Date     17 August 1978 ------------------------------------

                  Ogyrides limicola (D)                       5             1.7       0.6              30                 1.0
                  Paraprionospio pinnata (P)                  3             1.0       1.7              18                 2.0
                  Callianassa atla-ntica (D)                  2             0.7       1.2              12                 5.0
                  Glycera americana (P)                       2             0.7       0.6              12                 5.0
                  Glycinde solitaria (P)                      2             0.7       0.6              12                 5.0
                  Aricidea fragilis (P)                       2             0.7       1.2              12                 5.0
                  Clymenella torquata (P)                     2             0.7       0.6              12                 5.0
                  Stomatopoda (undet.)                        1             0.3       0.6                6                10.5
                  Pelecypoda (undet.) (M)                     1             0.3       0.6                6                10.5
                  Scoloplos sp. (P)                           1             0.3       0.6                6                10.5
                  Lumbrineris tenuis (P)                      1             0.3       0.6                6                10.5
                  Notomastus sp. (P)                          1             0.3       0.6                6                10.5
                  Asychis carolinae (P)                       1             0.3       0.6                6                10.5

                                                                         33

                 Table 18. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB04. (A = Amphipoda;
                            D = Decapoda; H    Hemichordata; M = Molluscs; 0 = Ophiuroidea; S = Stomatopoda; P = Polychaeta).

                                                                      SB04
                                                                                          2                               Rank
                                                           Total           Number/0.05m             Estimated              by
                              Species                     Number             -9        SD           Number/m2            Number

                 -----------------------------------   Collection Date    30 August 1977  -----------------------------------

                 Scoloplos rubra (P)                          7              2.3       1.5              42                 1.0
                 Ogyrides limicola (D)                        4              1.3       1.5              24                 3.0
                 Lumbrineris tenuis (P)                       4              1.3       1.2              24                 3.0
                 Glycinde solitaria (P)                       4              1.3       0.6              24                 3.0
                 Glycera americana (P)                        3              1.0       0.0              18                 5.5
                 Aricidea fragilis (P)                        3              1.0       1.7              18                 5.5
                 Squilla empusa (S)                           1              0.3       0.6               6                 9.5
                 Hemipholis elongata (0)                      1              0.3       0.6               6                 9.5
                 Anadara ovalis (M)                           1              0.3       0.6               6                 9.5
                 Diplodonta sp. (M)                           1              0.3       0.6               6                 9.5
                 Nereis succinea (P)                          1              0.3       0.6               6                 9.5
                 -5rilone@reismagna (P)                       1              0.3       0.6               6                 9.5

                 -----------------------------------   Collection Date    16 January   1978  ---------------------------------

                 Paraprionospio pinnata (P)                 17               5.7       2.1             103                 1.0
                 Heteromastus filiformis (P)                  5              1.7       0.6              30                 2.0
                 Ogyrides limfc-ola (D)                       3              1.0       0.0              18                 3.0
                 Lumbrineris tenuis (P)                       2              0.7       0.6              12                 4.0
                 Batea cathar-inensis (A)                     1              0.3       0.6               6                 9.0
                 Zaprella equilibra (A)                       1              0.3       0.6               6                 9.0
                 �accoglossus k@-w-alevskii (H)               1              0.3       0.6               6                 9.0
                 -Rulinia lateralis (M)                       1              0.3       0.6               6                 9.0
                 Scoloplos rubra (P)                          1              0.3       0.6               6                 9.0
                 Arabella iricolor (P)                        1              0.3       0.6               6                 9.0
                 Drilonereis magna (P)                        1              0.3       0.6               6                 9.0
                 Eteone heteropoda (P)                        1              0.3       0.6               6                 9.0
                 Haploscoloplos robustos (P)                  1              0.3       0.6               6                 9.0

                 ------------------------------------   Collection Date    24 April   1978 ----------------------------------

                 Ampelisca vadorum (A)                      12               4.0       1.7              73                 1.0
                 Lumbrineris tenuis (P)                     10               3.3       3.5              61                 2.0
                 Paraprionospio-pinnata    (P)                9              3.0       1.0              55                 3.0
                 Ogyrides limicola (D)                        2              0.7       0.6              12                 5.0
                 Tellinidae (undet.) (M)                      2              0.7       0.6              12                 5.0
                 Drilonereis magna (P)                        2              0.7       0.6              12                 5.0
                 Pelecypoda (undet. ) (M)                     1              0.3       0.6               6                10.5
                 Mulinia lateralis (M)                        1              0.3       0.6               6                10.5
                 Acteocina canaliculata (M)                   1              0.3       0.6               6                10.5
                 Glycera americana (P)                        1              0.3       0.6               6                10.5
                 Heteromastus filiformis (P)                  1              0.3       0.6               6                10.5
                 Glycinde solitaria (P)                       1              0.3       0.6               6                10.5
                 Aricidea fragilis (P)                        1              0.3       0.6               6                10.5
                 Haploscoloplos.sp. (P)                       1              0.3       0.6               6                10.5

                 -----------------------------------   Collection Date    17 August 1978  ----------------------------------

                 Ogyrides limicola (D)                      36             12.0        3.0             218                 1.0
                 Paraprionospio.pinnata (P)                 10               3.3       3.2              61                 2.0
                 Haploscoloplos fragilis (P)                  6              2.0       1.7              36                 3.0
                 Lumbrineris tenuis (P)                       3              1.0       1.0              18                 4.0
                 @iricidea fragilis (P)                       2              0.7       0.6              12                 5.0
                 Tellinidae (undet.) (M)                      1              0.3       0.6               6                 7.5
                 Glycera americans (P)                        1              0.3       0.6               6                 7.5
                 Glycinde solitaria (P)                       1              0.3       0.6               6                 7.5
                 Diopatra cuprea (P)                          1              0.3       0.6               6                 7.5

                                                                           34

                 Table 19. Macroinvertebrate species collected from the Sewee Bay channel station, SB05. (A
                            Amphipoda; C = Cumacea; D = Decapoda; I - Isopoda; M = Mollusca; 0 = Ophiuroidea;
                            P = Polychaeta).

                                                                        SB05
                                                                                                                           Rank
                                                          Total             Number/0.05m2           Estimated               by
                            Species                      Number             'i         SD           Number/m2            Number

                 -----------------------------------   Collection Date      23 August  1977 --------------------------------------

                 Paraprionospio pinnata (P)                  27             9.0        2.7               164                1.0
                 lumbrineris ten@u-is(P)                      5             1.7        0.6               30                 2.0
                 Glycera americana (P)                        3             .1.0       1.0               18                 3.0
                 Hemipholis elo      a (0)                    2             0.7        0.6               12                 4.5
                 Scoloplos rubra (P)                          2             0.7        0.6               12                 4.5
                 Ogyrides limicola (D)                        1             0.3        0.6                 6                8.5
                 Polinices dglicatus     (M)                  1             0.3        0.6                 6                8.5
                 Mulinia lateralis (M)                        1             0.3        0.6                 6                8.5
                 Ilyanassa obsoleta (M)                       1             0.3        0.6                 6                8.5
                 -dlycinde s-olitaria (P)                     1             0.3        0.6                 6                8.5
                 Aricidea fragilis (P)                        1             0.3        0.6                 6                8.5

                 -----------------------------------   Collection Date      30 January 1978 -------------------------------------

                 Mulinia lateralis (M)                        7             2.3        1.5               42                 1.0
                 6gyrides limicola (D)                        3             1.0        1.0               12                 2.0
                 Leucon americanus; (C)                       2             0.6        0.6               12                 3.5
                 Nereis succinea (P)                          2             0.6        1.2               12                 3.5
                 Palaemonetes vulgaris   (D)                  1             0.3        0.6                 6                9.0
                 Callinectes ornatus (D)                      1             0.3        0.6                 6                9.0
                 Caprella equilibra (A)                       1             0.3        0.6                 6                9.0
                 Mercenaria mercenaria (M)                    1             0.3        0.6                 6                9.0
                 Abra aegualis (M)                            1             0.3        0.6                 6                9.0
                 Pectinaria gouldii (P)                       1             0.3        0.6                 6                9.0
                 _ffeteromastus filiformis (P)                1             0.3        0.6                 6                9.0
                 Arabella iricolor (P)                        1             0.3        0.6                 6                9.0
                 Paraprionospio pinnata (P)                   1             0.3        0.6                 6                9.0

                 ------------------------------------    Collection Date    28 April   1978 --------------------------------------

                 Haploscoloplos sp. (P)                      10             3.3        4.9               61                 1.0
                 Mulinia lateralis (M)                        8             2.7        2.9               48                 2.0
                 Ogyrides limicola (D)                        4             1.3        1.2               24                 3.0
                 Heteromastus filiformis (P)                  3             1.0        1.0               18                 4.5
                 Paraprionospio pinnata (P)                   3             1.0        1.7               18                 4.5
                 Teucon americanus (C)                        2             0.7        0.6               12                 6.5
                 Glycera americana (P)                        2             0.7        0.6               12                 6.5
                 Ampelisca vadorum (A)                        1             0.3        0.6                 6                9.5
                 Edotea montosa (f)                           1             0.3        0.6                 6                9.5
                 Tolinices duplicatus    (M)                  1             0.3        0.6                 6                9.5
                 Tharyx marioni (P)                           1             0.3        0.6                 6                9.5

                 ------------------------------------    Collection Date    3 August   1978 -------------------------------------

                 Ogyrides limicola (D)                        3             1.0        1.7               18                 1.0
                 Zlycera am-ericana (P)                       2             0.7        0.6               12                 3.0
                 Heteromastus filiformis (P)                  2             0.7        0.6               12                 3.0
                 -@araprionospio pinnata (P)                  2             0.7        1.2               12                 3.0
                 Lumbrineris tenuis (P)                       1             0.3        0.6                 6                5.5
                 -Brilonereis magna (P)                       1             0.3        0.6                 6                5.5

                                                                          35

                  Table 20. Macroinvertebrate species collected from the Sewee Bay intertidal station, SB06. (I
                             Isopoda; M = Mollusca; P = Polychaeta).

                                                                        SB06
                                                                                                                             Rank
                                                             Total            Number/0.05m2           Estimated              by
                              Species                        Number            'K         SD          Number/m2            Number

                  ------------------------------------   Collection Date    30 August 1977  ------------------------------------

                  Lumbrineris tenuis (P)                        12            4.0         1.0             73                 1.0
                  'Nereis succinea (P)                           7            2.3         1.2             42                 2.0
                  Aricidea fragilis (P)                          4            1.3         0.6             24                 3.0
                  Drilonereis magna (P)                          2            0.7         0.6             12                 4.0
                  Cyathura burbanki (1)                          1            0.3         0.6               6                6.5
                  Mulinia lateralis (M)                          1            0.3         0.6               6                6.5
                  Ilyanassa obsoleta (M)                         1            0.3         0.6               6                6.5
                  Glycera americana (P)                          1            0.3         0.6               6                6.5

                  -------------------------------------  Collection Date    30 January 1978 ----------------------------------

                  Lumbrineris tenuis (P)                         6            2.0         2.0             36                 1.0
                  Ilyanassa obsoleta (M)                         4            1.3         1.5             24                 2.5
                  Heteromastus filiformis (P)                    4            1.3         0.6             24                 2.5
                  Glycera americana (P)                          2            0.7         1.2             12                 5.0
                  Aricidea fragilis (P)                          2            0.7         0.6             12                 5.0
                  Haploscoloplos sp. (P)                         2            0.7         0.6             12                 5.0
                  Cyathura polita (1)                            1            0.3         0.6               6                8.5
                  Mercenaria mercenaria   (M)                    1            0.3         0.6               6                8.5
                  Mulinia lateralis (M)                          1            0.3         0.6               6                8.5
                  Nereis succinea (P)                            1            0.3         0.6               6                8.5

                  -------------------------------------   Collection Date    25 April 1978  -----------------------------------

                  Haploscoloplos fragilis (P)                   27            9.0        10.4             164                1.0
                  Lumbrineris tenuis (P)                        17            5.7         6.4             103                2.0
                  Aricidea fragilis (P)                          8            2.7         2.3             48                 3.0
                  Streblospio benedicti   (P)                    3            1.0         1.0             18                 4.0
                  Glycera americana (P)                          2            0.7         0.6             12                 5.5
                  Mediomastus californiensis (P)                 2            0.7         1.2             12                 5.5
                  Edotea montosa (1)                             1            0.3         0.6               6                8.0
                  Heteromastus filiformis (P)                    1            0.3         0.6               6                8.0
                  Nereis succinea (P)                            1            0.3         0.6               6                8.0

                  -------------------------------------   Collection Date    24 July 1978 ------------------------------------

                  Mulinia lateralis (M)                          1            0.3         0.6               6                3.0
                  Glycera americana (P)                          1            0.3         0.6               6                3.0
                  Heteromastus filiformis (P)                    1            0.3         0.6               6                3.0
                  Drilonereis magna (P)                          1            0.0         0.0               6                3.0
                  Aricidea fragilis (P)                          1            0.3         0.6               6                3.0

                                                                         36

                  Table 21. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB07. (A
                           Amphipoda; C = Cumacea; D = Decapoda; I = Isopoda; M = Mollusca; 0 = Ophiuroidea;
                           P = Polychaeta; S = Stomatopoda).

                                                                       SB07
                                                                                                                              Rank
                                                             Total             Number/0.05m  2         Estimated               by
                              Species                       Number             3@        SD            Number/m2             Number

                  ------------------------------------  Collection  Date    25 August 1977   ------------------------------------

                  Nereis succinea (P)                            7             2.3       2.1               42                  1.0
                  Glycera americana (P)                          4             1.3       1.5               24                  2.0
                  Lumbrineris tenuis (P)                         3             1.0       1.0               18                  3.0
                  Scoloplos rubra (P)                            2             0.7       1.2               12                  4.5
                  Tharyx marioni (P)                             2             0.7       0.6               12                  4.5
                  Alpheus normanni (D)                           1             0.3       0.6                 6                 9.5
                  Batea catharinensis     (A)                    1             0.3       0.6                 6                 9.5
                  Hemipholis elongata     (0)                    1             0.3       0.6                 6                 9.5
                  Zrepidula fornicata     (M)                    1             0.3       0.6                 6                 9.5
                  Notomastus lobatus (P)                         1             0.3       0.6                 6                 9.5
                  Glycinde solitaria (P)                         1             0.3       0.6                 6                 9.5
                  Arabella iricolor (P)                          1             0.3       0.6                 6                 9.5
                  Drilonereis magna (P)                          1             0.3       0.6                 6                 9.5

                  ------------------------------------   Collection Date    30 January   1978 -----------------------------------

                  Mulinia lateralis (M)                        10              3.3       3.5               61                  1.0
                  Nereis succinea (P)                            5             1.7       1.5               30                  2.0
                  Paraprionos2io pinnata (P)                     4             1.3       1.5               24                  3.0
                  Glycera americans (P)                          3             1.0       1.0               18                  4.0
                  Xanthidae (undet.) (D)                         2             0.7       1.2               12                  7.0
                  Ampelisca vadorum (A)                          2             0.7       0.6               12                  7.0
                  Cyathura polita (1)                            2             0.7       1.2               12                  7.0
                  Streblospio benedicti   (P)                    2             0.7       1.2               12                  7.0
                  Cirratulidae (undet.)   (P)                    2             0.7       1.2               12                  7.0
                  Palaemonetes vulgaris   (D)                    1             0.3       0.6                 6                 18.5
                  Upogebia affinis (D)                           1             0.3       0.6                 6                 18.5
                  Gammaridae (undet.) (A)                        1             0.3       0.6                 6                 18.5
                  Melitidae (undet.) (A)                         1             0.3       0.6                 6                 18.5
                  Nemertina sp. A                                1             0.3       0.6                 6                 18.5
                  Nemertina sp. B                                1             0.3       0.6                 6                 18.5
                  Mitrella lunata   (M)                          1             0.3       0.6                 6                 18.5
                  Macoma sp. (M)                                 1             0.3       0.6                 6                 18.5
                  Lumbrineris tenuis      (P)                    1             0.3       0.6                 6                 18.5
                  Glycinde solitaria      (P)                    1             0.3       0.6                 6                 18.5
                  Marphysa sanguinea      (P)                    1             0.3       0.6                 6                 18.5
                  Arabella -iricolor      (P)                    1             0.3       0.6                 6                 18.5
                  Haploscoloplos fragilis (P)                    1             0.3       0.6                 6                 18.5
                  Oligochaeta (undet.)                           1             0.3       0.6                 6                 18.5
                  Aricidea fragilis (P)                          1             0.3       0.6                 6                 18.5
                  Lysidice ninetta (P)                           1             0.3       0.6                 6                 18.5
                  Subadyte pellucida (P)                         1             0.3       0.6                 6                 18.5
                  Pholoe minuta (P)                              1             0.3       0.6                 6                 18.5

                  --7 ----------------------------------  Collection Date      24 April 1978 ------------------------------------

                  Ampelisca vadorum (A)                        12              4.0       2.0               73                  1.5
                  Lumbrineris tenuis (P)                       12              4.0       2.0               73                  1.5
                  Oxyurostylis smithi (C)                      11              3.7       4.0               67                  3.0
                  Glycera americana (P)                          4             1.3       1.2               24                  4.5
                  Nereis succinea (P)                            4             1.3       0.6               24                  4.5
                  Drilonereis magna (P)                          3             1.0       1.0               18                  6.0
                  Xanthidae (undet.)      (D)                    2             0.7       0.6               12                  8.5
                  Gammaridae (undet.)(A)                         2             0.7       1.2               12                  8.5
                  Nemertina (undet.)                             2             0.7       0.6               12                  8.5
                  Haploscoloplos sp.      (P)                    2             0.7       0.6               12                  8.5
                  Pagurus longicarpus (D)                        1             0.3       0.6                 6                 17.5
                  Squilla empusa (S)                             1             0.3       0.6                 6                 17.5
                  Caprella equilibra      (A)                    1             0.3       0.6                 6                 17.5
                  Nucula proxima (M)                             1             0.3       0.6                 6                 17.5
                  Mercenaria mercenaria (M)                      1             0.3       0.6                 6                 17.5
                  Abra aegualis (M)                              1             0.3       0.6                 6                 17.5
                  Pelecypoda (undet.) (M)                        1             0.3       0.6                 6                 17.5
                  Acteocina canaliculata (M)                     1             0.3       0.6                 6                 17.5
                  Tellinidae (undet.) (M)                        1             0.3       0.6                 6                 17.5

                                                                        37

                   Table 21. (Cont.

                                                                           SB07
                                                                                                                                Rank
                                                                Total            Number/0.05m2            Estimatel              by
                               Species                          Number            -9        SD            Number/m            Number

                   Scoloplos rubra (P)                             1             0.3        0.6                6                17.5
                   Pectinaria gouldii (P)                          1             0.3        0.6                6                17.5
                   Scoloplos sp. (P)                               1             0.3        0.6                6                17.5
                   Paraprionospio pinnata   (P)                    1             0.3        0.6                6                17.5
                   Mediomastus californiensis    (P)               1             0.3        0.6                6                17.5

                   -------------------------------------   Collection Date     17 August 1978  ------------------------------------

                   Lumbrineris tenuis (P)                          15            5.0        1.0               91                 1.0
                   Panopeus herbstii (D)                           6             2.0        1.0               36                 2.0
                   Podarke obscura (P)                             4             1.3        1.2               24                 3.0
                   Cyathura buri@anki (1)                          3             1.0        1.0               18                 5.0
                   Heteromastus filiformis (P)                     3             1.0        1.0               18                 5.0
                   Aricidea fragilis (P)                           3             1.0        1.0               18                 5.0
                   Melita sp. (A)                                  2             0.7        0.6               12                 9.0
                   Drilonereis magna (P)                           2             0.7        0.6               12                 9.0
                   Lepidonotus sublevis    (P)                     2             0.7        0.6               12                 9.0
                   Diplocirrus capensis    (P)                     2             0.7        0.6               12                 9.0
                   Cirratulidae (undet.).(P)                       2             0.7        1.2               12                 9.0
                   Alpheus normanni (D)                            1             0.3        0.6                6                15.5
                   Nucula proxima (M)                              1             0.3        0.6                6                15.5
                   Glycera americana (P)                           1             0.3        0.6                6                15.5
                   Scoloplos rubra (P)                             1             0.3        0.6                6                15.5
                   Nereis succinea (P)                             1             0.3        0.6                6                15.5
                   Glycinde solitaria (P)                          1             0.3        0.6                6                15.5
                   Arabella iricolor (P)                           1             0.3        0.6                6                15.5
                   Syllis spongicola (P)                           1             0.3        0.6                6                15.5

                                                                          38

                   Table 22. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB08. (A
                             Amphipoda; C = Cumacea; D = Decapoda; H = Hemichordata; I = Isopoda; M = Mollusca;
                             0 = Ophiuroidea; P = Polychaeta).

                                                                       SBO8
                                                                                            2                                 Rank
                                                             Total             Number/0.05m            Estimated              by
                                Species                     Number             -x        SD            Number/m2            Number

                   -----------------------------------  Collection  Date     25 August 1977 ------------------------------------

                   Lumbrineris tenuis (P)                        6             2.0       2.0                36                1.5
                   Paraprionospio pinnata (P)                    6             2.0       2.7                36                1.5
                   Glycera americana (P)                         3             1.0       0.0                18                3.5
                   Scoloplos rubra (P)                           3             1.0       1.0                18                3.5
                   Cyathura burbanki (1)                         1             0.3       0.6                6                 7.0
                   Hemipholis elongata (0)                       1             0.3       0.6                6                 7.0
                   Saccoglossus kowalevskii (H)                  1             0.3       0.6                6                 7.0
                   Gly@,@ solitaria (P)                          1             0.3       0.6                6                 7.0
                   Drilonereis magna (P)                         1             0.3       0.6                6                 7.0

                   -----------------------------------  Collection Date      30 January 1978 ----------------------------------

                   Paraprionospio pinnata (P)                    11            3.7       4.7                67                1.0
                   Mulinia lateralis                             7             2.3       2.5                42                2.0
                   Lumbrineris tenuis (P)                        6             2.0       1.7                36                3.0
                   Nereis succinea (P)                           5             1.7       0.6                30                4.5
                   Haploscoloplos robustos (P)                   5             1.7       0.6                30                4.5
                   Heteromastus filiformis (P)                   4             1.3       1.2                24                6.0
                   Ogyrides limicola (D)                         3             1.0       1.7                18                7.5
                   Mitrella lunata (M)                           3             1.0       1.0                18                7.5
                   Pinnixa sp. (D)                               2             0.7       1.2                12                11.0
                   Ampelisca vadorum (A)                         2             0.7       0.6                12                11.0
                   Cyathura burbanki (1)                         2             0.7       1.2                12                11.0
                   Tellinidae (undet.) (M)                       2             0.7       1.2                12                11.0
                   Lysidice ninetta (P)                          2             0.7       1.2                12                11.0
                   Mercenaria mercenaria (M)                     1             0.3       0.6                6                 16.0
                   Glycera americana (P)                         1             0.3       0.6                6                 16.0
                   Glycinde solitaria (P)                        1             0.3       0.6                6                 16.0
                   Oligochaeta (undet.)                          1             0.3       0.6                6                 16.0
                   Polydora sp. (P)                              1             0.3       0.6                6                 16.0

                   ------------------------------------   Collection Date      24 April 1978 -----------------------------------

                   Haploscoloplos fragilis (P)                   36            12.0      9.0              218                 1.0
                   Ampelisca vadorum (A)                         12            4.0       1.0                73                2.0
                   Spionidae (undet.) (P)                        6             2.0       2.0                36                3.0
                   Heteromastus filiformis (P)                   5             1.7       2.1                30                4.5
                   Lunbrineris tenuis (P)                        5             1.7       0.6                30                4.5
                   Tellinidae (undet.) (M)                       4             1.3       1.2                24                6.0
                   Melita nitida (A)                             3             1.0       1.7                18                8.0
                   Nereis succinea (P)                           3             1.0       1.7                18                8.0
                   GlyS_@ solitaria (P)                          3             1.0       1.0                18                8.0
                   Leucon americanus (C)                         2             0.7       0.6                12                10.0
                   Xanthidae (undet.) (D)                        1             0.3       0.6                6                 14.5
                   Nucula proxima (M)                            1             0.3       0.6                6                 14.5
                   Pelecypoda (undet.) (M)                       1             0.3       0.6                6                 14.5
                   Glycera americana (P)                         1             0.3       0.6                6                 14.5
                   Tharyx marioni (P)                            1             0.3       0.6                6                 14.5
                   Drilonereis magna (P)                         1             0.3       0.6                6                 14.5
                   Eteone heteropoda (P)                         1             0.3       0.6                6                 14.5
                   Tharyx setigera (P)                           1             0.3       0.6                6                 14.5

                   ------------------------------------   Collection Date      3 August 1978 -----------------------------------

                   Ogyrides limicola (D)                         33            11.0      8.0              200                 1.0
                   Haploscoloplos fragilis (P)                   6             2.0       2.7                36                2.0
                   Lurrbrineris tenuis (P)                       5             1.7       1.2                30                3.0
                   Paraprionospio pinnata (P)                    3             1.0       1.7                18                4.0
                   Glycera americana (P)                         1             0.3       0.6                6                 5.5
                   Not3mastus sp.                                1             0.3       0.6                6                 5.5

                                                                        39

                    Table 23. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB09. (A = Amphipoda;
                               H = Hemichordata; I = Isopoda; M = Mollusca; P = Polychaeta; Py = Pycnogonida).

                                                                               SB09
                                                                                                                                          Rank
                                                                    Total               Number/0.05m2            Estimated                by
                                  Species                          Number               x         SD             Number/M2             Number

                    ------------------------------------       Collection Date     23 August 1977    ------------------------------------

                    Lumbrineris tenuis (P)                             49               16.3      11.6              297                   1.0
                    Paraprionospio pinnata (P)                         26               8.7       14.2              158                   2.0
                    Aricidea fragilis (P)                              14               4.7       3.1                   85                3.0
                    Sacoglossus kowalevskii (H)                        6                2.0       1.7                   36                4.0
                    Mulinia lateralis (M)                              5                1.7       1.2                   30                5.5
                    Glycinde solitaria (P)                             5                1.7       1.5                   30                5.5
                    Glycera americana (P)                              3                1.0       1.0                   18                7.0
                    Heteromastus filiformis (P)                        2                0.7       1.2                   12                9.0
                    Haploscoloplos fragilis (P)                        2                0.7       0.6                   12                9.0
                    Haploscoloplos foliosus (P)                        2                0.7       1.2                   12                9.0

                    ------------------------------------       Collection Date      31  January   1978 -----------------------------------

                    Mulinia lateralis (M)                              74               24.7      11.9              449                   1.0
                    Lumbrineris tenuis (P)                             19               6.3       9.3               115                   2.0
                    Edotea montosa (1)                                 5                1.7       2.9                   30                4.5
                    Odostomia sp. (P)                                  5                1.7       2.9                   30                4.5
                    Heteromastus filiformis (P)                        5                1.7       2.9                   30                4.5
                    Haploscoloplos sp. (P)                             5                1.7       2.1                   30                4.5
                    Nereis succinea (P)                                4                1.3       2.3                   24                7.0
                    Glycera americana (P)                              3                1.0       1.0                   18                8.0
                    Tellinidae -(undet.) (M)                           2                0.7       0.6                   12                10.5
                    Eteone heteropoda (P)                              2                0.7       0.6                   12                10.5
                    Mediomastus californiensis (P)                     2                0.7       0.6                   12                10.5
                    Aricidea fragilis (P)                              2                0.7       1.2                   12                10.5
                    Paraphoxus spinosus (A)                            1                0.3       0.6                   6                 20.0
                    Gammarus mucronatus (A)                            1                0.3       0.6                   6                 20.0
                    Tanystylum calicirostre     (PY)                   1                0.3       0.6                   6                 20.0
                    Actinaria (undet.)                                 1                0.3       0.6                   6                 20.0
                    Polinices lacteus (M)                              1                0.3       0.6                   6                 20.0
                    Mercenaria mercenaria     (M)                      1                0.3       0.6                   6                 20.0
                    Acteocina canaliculata (M)                         1                0.3       0.6                   6                 20.0
                    Streblospio benedicti (P)                          1                0.3       0.6                   6                 20.0
                    Marphysa sanguinea (P)                             1                0.3       0.6                   6                 20.0
                    Drilonereis magna (P)                              1                0.3       0.6                   6                 20.0
                    Leptdonotus sublevis (P)                           1                0.3       0.6                   6                 20.0
                    Polydora ligni (P)                                 1                0.3       0.6                   6                 20.0
                    Paraprionospio pinnata (P)                         1                0.3       0.6                   6                 20.0
                    Thelepus setosus (P)                               1                0.3       0.6                   6                 20.0
                    Nereinides unidentata (P)                          1                0.3       0.6                   6                 20.0

                    -------------------------------------      Collection Date      25 April 1978    --------------------------------------

                    Lumbrineris tenuis (P)                             51               17.0      10.0              309                   1.0
                    Haploscoloplos fragilis (P)                        21               7.0       4.0               127                   2.0
                    Heteromastus filiformis (P)                        16               5.3       4.5                   97                3.0
                    Streblospio benedicti (P)                          12               4.0       5.3                   73                4.5
                    Cirratulidae (undet.) (P)                          12               4.0       4.6                   73                4.5
                    Tellinidae (undet.) (M)                            7                2.3       3.2                   42                6.0
                    Aricidea fragilis (P)                              5                1.7       0.6                   30                7.0
                    Glycera americana (P)                              3                1.0       1.0                   18                8.0
                    Glycinde solitaria (P)                             2                0.7       1.2                   12                10.0
                    Oligochaeta (undet.)                               2                0.7       1.2                   12                10.0
                    Clymenella torquata (P)                            2                0.7       1.2                   12                10.0
                    Ampelisca vadorum (A)                              1                0.3       0.6                   6                 13.5
                    Edotea montosa (1)                                 1                0.3       0.6                   6                 13.5
                    Nemertina (undet.)                                 1                0.3       0.6                   6                 13.5
                    Eteone heteropoda (P)                              1                0.3       0.6                   6                 13.5

                    -------------------------------------      Collection Date      24 July 1978  --------------------------------------

                    Lumbrineris tennis (P)                             12               4.0       3.0                   73                1.0
                    Glycera americana (P)                              3                1.0       1.0                   18                2.5
                    Aricidea fragilis (P)                              3                1.0       1.0                   18                2.5
                    Cyathura burbanki (1)                              1                0.3       0.6                   6                 5.5
                    Nemertina (undet.)                                 1                0.3       0.6                   6                 5.5
                    Mulinia lateralis (M)                              1                0.3       0.6                   6                 5.5
                    Glycinde solitaria (P)                             1                0.3       0.6                   6                 5.5

                  Table24,. Macroinvertebrate species collected from the Sewee Bay channel station, SB10. (A = Amphipoda;
                            C = Cumacea; D = Decapoda; H = Hemicordata; M = Mollusca; P = Polychaeta).

                                                                        SB10
                                                                                             2                                  Rank
                                                             Total              Number/0.05m            Estimate5               by
                             Species                         Number             -:w       SD           Number/m              Number

                  ------------------------------------   Collection  Date   23 August 1977   ------------------------------------

                  Paraprionospio pinnata (P)                      43            14.3     11.9                  261              1.0
                  Lumbrineris tenuis (P)                          10            3.3       1.2                  61               2.0
                  Ogyrides limicola (D)                           8             2.7       1.2                  48               3.0
                  Notomastus lobatus (P)                          5             1.7       1.5                  30               4.0
                  Caprella equilibra (A)                          3             1.0       1.7                  18               5.5
                  Glycinde solitaria (P)                          3             1.0       1.0                  18               5.5
                  Batea catharinensis (A)                         2             0.7       1.2                  12               8.0
                  Glycera americana (P)                           2             0.7       0.6                  12               8.0
                  Tharyx.annulosus (P)                            2             0.7       1.2                  12               8.0
                  Alpheus heterochaelis   (D)                     1             0.3       0.6                  6                11.5
                  Ampelisca vadorum (A)                           1             0.3       0.6                  6                11.5
                  Sacoglossus kowalevskii (H)                     1             0.3       0.6                  6                11.5
                  Maldanidae (undet.) (P)                         1             0.3       0.6                  6                11.5

                  ------------------------------------   Collection Date     30 January  1978 -----------------------------------

                  Ogyrides limicola (D)                           8             2.7       1.5                  48               1.0
                  Heteromastus.filiformis (P)                     2             0.7       1.2                  12               2.5
                  Paraprionospio pinnata (P)                      2             0.7       0.6                  12               2.5
                  Leucon americanus (C)                           1             0.3       0.6                  6                4.5
                  Abra aegualis (M)                               1             0.3       0.6                  6                4.5

                  -------------------------------------   Collection Date       28 April 1978 ------------------------------------

                  Heteromastus filiformis (P)                     67            22.3      5.5                  406              1.0
                  Paraprionospio pinnata (P)                      20            6.7       1.5                  121              2.0
                  Gemma gemma (M)                                 12            4.0       6.1                  73               3.5
                  Nereis succinea (P)                             12            4.0       3.6                  73               3.5
                  Haploscoloplos fragilis .(P)                    11            3.7       3.2                  67               5.0
                  Ogyrides limicola (D)                           9             3.0       3.6                  55               6.0
                  Lumbrineris tenuis (P)                          6             2.0       2.0                  36               8.5
                  @rilonereis magna (P)                           6             2.0       1.0                  36               8.5
                  Clymenella torquata (P)                         6             2.0       2.0                  36               8.5
                  Streblospio benedicti (P)                       6             2.0       2.0                  36               8.5
                  Tellinidae (undet.) (M)                         5             1.7       1.5                  30               11.0
                  Mulinia lateralis (M)                           4             1.3       0.6                  24               12.5
                  Nereidae (undet.) (P)                           4             1.3       1.5                  24               12.5
                  Ampelisca vadorum (A)                           3             1.0       0.0                  18               15.5
                  Nemertina (undet.)                              3             1.0       1.0                  18               15.5
                  Haploscoloplos fragilis (P)                     3             1.0       0.0                  18               15.5
                  Aricidea fragilis (P)                           3             1.0       0.0                  18               15.5
                  Oxyurostylis smithi (C)                         2             0.7       1.2                  12               19.0
                  Polinices lacteus (M)                           2             0.7       0.6                  12               19.0
                  Mediomastus californiensis (P)                  2             0.7       1.2                  12               19.0
                  Solen viridis (M)                               1             0.3       0.6                  6                24.0
                  Abra aequalis (M)                               1             0.3       0.6                  6                24.0
                  Pelecypoda. (undet.) (M)                        1             0.3       0.6                  6                24.0
                  Tellinidae (undet.) (M)                         1             0.3       0.6                  6                24.0
                  Glycera americana (P)                           1             0.3       0.6                  6                24.0
                  Pectinaria gouldii (P)                          1             0.3       0.6                  6                24.0
                  Cirratulidae (undet.) (P)                       1             0.3       0.6                  6                24.0

                  -------------------------------------   Collection Date       3 August 1978 -----------------------------------

                  Ogyrides limicola (D)                           64            21.3      4.0                  388              1.0
                  Heteromastus filiformis (P)                     24            8.0       3.6                  145              2.0
                  Haploscoloplo's fragilis (P)                    19            6.3       3.1                  115              3.5
                  Paraprionospio pinnata (P)                      19            6.3       6.8                  115              3.5
                  Glycera. americans (P)                          2             0.7       0.6                  12               6.0
                  Nereis succinea (P)                             2             0.7       1.2                  12               6.0
                  Glycfnde solitaria (P)                          2             0.7       1.2                  12               6.0
                  Pinnixa sayana (D)                              1             0.3       0.6                  6                10.0
                  Sigambra tentaculata (P)                        1             0.3       0.6                  6                10.0
                  Lumbrineris tenuis (P)                          1             0.3       0.6                  6                10.0
                  Diopatra cuprea (P)                             1             0.3       0.6                  6                10.0
                  Oligochaeta (undet.)                            1             0.3       0.6                  6                10.0

                                                                        41

                     Table25. Macroinvertebrate species collected from the Sewee Bay intertidal station, SB11. (M
                               Mollusca; P = Polychaeta).

                                                                           SBII
                                                                                                                             Rank
                                                             Total            Number/0.05m  2          Estimated              by
                               Species                       Number            -K        SD            Number/m2            Number

                     -----------------------------------   Collection Date    30 August 1977 ----------------------------------

                     Heteromastus filiformis (P)               91             30.3      13.8              552                 1.0
                     Nereis succinea (P)                       37             12.3      10.4              224                 2.0
                     Gemma gemma (M)                           11              3.7       1.5                67                3.0
                     Aricidea fragilis (P)                     10              3.3       4.9                61                4.0
                     Lumbrineris tenuis (P)                      6             2.0       3.5                36                5.0
                     Laeonereis culveri (P)                      3             1.0       1.0                18                6.0
                     Glycera americana (P)                       1             0.3       0.6                 6                8.0
                     Diopatra cuprea (P)                         1             0.3       0.6                 6                8.0
                     Drilonereis magna (P)                       1             0.3       0.6                 6                8.0

                     ----------------------------------  Collection Date     10 February 1978  ---------------------------------

                     Heteromastus filiformis (P)               47             15.7      10.1              285                 1.0
                     Gemma gemma (M)                           42             14.0      13.2              255                 2.0
                     Paraprionospio pinnata (P)                16              5.3       5.5                97                3.0
                     Nereidae (undet.) (P)                       5             1.7       1.2                30                4.0
                     Lumbrineris tenuis (P)                      2             0.7       1.2                12                5.0
                     Mercenaria mercenaria   (M)                 1             0.3       0.6                 6                8.5
                     Tagelus plebeius (P)                        1             0.3       0.6                 6                8.5
                     Glycera americana (P)                       1             0.3       0.6                 6                8.5
                     Nereis succinea (P)                         1             0.3       0.6                 6                8.5
                     Diopatra cuprea (P)                         1             0.3       0.6                 6                8.5
                     Haploscoloplos sp. (P)                      1             0.3       0.6                 6                8.5

                     ------------------------------------   Collection Date    28 April  1978 ----------------------------------

                     Heteromastus filiformis (P)               51             17.0       6.1              309                 1.0
                     Gemma gemma. (M)                          12              4.0       6.1                73                2.5
                     Nereis succinea (P)                       12              4.0       3.6                73                2.5
                     Haploscoloplos fragilis (P)               11              3.7       3.2                67                4.0
                     Lumbrineris tenuis (P)                      6             2.0       2.0                36                5.5
                     Drilonereis magna (P)                       6             2.0       1.0                36                5.5
                     Nereidae (undet.) (P)                       4             1.3       1.5                24                7.0
                     Nemertina (undet.)                          3             1.0       1.0                18                8.5
                     Aricidea fragilis (P)                       3             1.0       0.0                18                8.5
                     Polinices lacteus (M)                       2             0.7       0.6                12               10.5
                     Streblospio benedicti   (P)                 2             0.7       1.2                12               10.5
                     Tellinidae (undet.) (M)                     1             0.3       0.6                 6               12.0

                     -----------------------------------   Collection Date    18 August  1978 ----------------------------------

                     Nereis succinea (P)                       48             16.0       8.7              291                 1.0
                     Heteromastus filiformis (P)               24              8.0       6.1              145                 2.0
                     Gemma gemma (M)                           19              6.3       1.5              115                 3.0
                     Haploscoloplos fragilis (P)               18              6.0       4.4              109                 4.0
                     Aricidea fragilis (P)                       3             1.0       1.0                18                5.0
                     Nemertina (undet.)                          1             0.3       0.6                 6                7.0
                     Lumbrineris tenuis (P)                      1             0.3       0.6                 6                7.0
                     Drilonereis magna (P)                       1             0.3       0.6                 6                7.0

                                                                          42

                  Table 26. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB12. (A = Amphipoda;
                            I = Isopoda; M = Mollusca; P = Polychaeta).

                                                                            SB12
                                                                                                                            Rank
                                                             Total            Number/0.05m2           Estimated              by
                             Species                         Number            T         SD           Number/M2             Number

                  ----------------------------------------   Collection Date  = 30 August 1977 --------------------------------

                  Haploscoloplos robustos (P)                  41             13.7       3.8              249                1.0
                  Lumbrineris tenuis (P)                       11              3.7       3.8               67                2.0
                  Heteromastus filiformis (P)                    9             3.0       2.7               55                3.0
                  Aricidea fragilis (P)                          8             2.7       3.1               48                4.0
                  Cyathura burbanki (1)                          1             0.3       0.6                 6               5.5
                  Streblospio benedic-ti  (P)                    1             0.3       0.6                 6               5.5

                  ----------------------------------------   Collection Date = 31  January 1978  -------------------------------

                  Nereis succinea (P)                          34             11.3       7.1              206                1.0
                  Heteromastus filiformis (P)                  18              6.0       3.0              109                2.0
                  Paraprionospio pinnata (P)                   13              4.3       4.0               79                3.0
                  Haploscoloplos fragilis (P)                    6             2.0       1.0               36                4.0
                  Glycera americana (P)                          3             1.0       1.0               18                5.5
                  Nereidae (undet.) (P)                          3             1.0       1.0               18                5.5
                  Mulinia lateralis (M)                          1             0.3       0.6                 6               8.0
                  Tellinidae (undet.) (M)                        1             0.3       0.6                 6               8.0
                  Oligochaeta (undet.)                           1             0.3       0.6                 6               8.0

                  -----------------------------------------   Collection Date  = 25 April 1978 --------------------------------

                  Haploscoloplos fragilis (P)                  21              7.3       9.3              127                1.0
                  Streblospio benedicti (P)                    18              5.0       8.7              109                2.0
                  Mediomastus californiensis   (P)             10              3.3       3.2               61                3.0
                  Heteromastus filiformis (P)                    8             2.7       1.2               48                4.0
                  Paraprionospio pinnata (P)                     4             1.3       1.5               24                5.0
                  Lumbrineris tenuis (P)                         3             1.0       1.7               18                6.0
                  Ampelisca vadorum (A)                          2             0.7       1.2               12                7.5
                  Glycera americana (P)                          2             0.7       0.6               12                7.5
                  Ogyrides limicola (D)                          1             0.3       0.6                 6              12.5
                  Ilyanassa obsoleta (M)                         1             0.3       0.6                 6              12.5
                  Tellinidae (undet.) (M)                        1             0.3       0.6                 6              1.2.5
                  Nereis succinea (P)                            1             0.3       0.6                 6              12.5
                  Eteone heteropoda (P)                          1             0.3       0.6                 6              12.5
                  Oligochaeta (undet.)                           1             0.3       0.6                 6              12.5
                  Aricidea fragilis (P)                          1             0.3       0.6                 6              12.5
                  Cirratulidae (undet.)   (P)                    1             0.3       0.6                 6              12.5

                  ----------------------------------------   Collection Date     18 August 1978 -------------------------------

                  Haploscoloplos fragilis (P)                  24              8.0       7.6              145               1.0
                  Heteromastus filiformis (P)                    8             2.7       1.5               48               2.0
                  Paraprionospio pinnata (P)                     5             1.7       1.5               30               3.0
                  Lumbrineris tenuis (P)                         3             1.0       0.0               18               4.0
                  Oligochaeta (undet.)                           2             0.7       0.6               12               5.0
                  Sigambra tentaculata (P)                       1             0.3       0.6                 6              7.0
                  Glycinde solitaria (P)                         1             0.3       0.6                 6              7.0
                  Aricidea fragilis (P)                          1             0.3       0.6                 6              7.0

                                                                        43

                    Table 27. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB13. (A =
                               Amphipoda; C = Cumacea; D = Decapoda; Ho @ Holothuroidea; I = Isopoda; M = Mollusca;
                               P = Polychaeta).

                                                                          SB13
                                                                                                                            Rank
                                                            Total            Number/0.05m2            Estimatel              by
                               Species                      Number            -9        SD            Number/m             Number

                    -----------------------------------   Collection Date    30 August 1977 ----------------------------------

                    Lumbrineris tenuis (P)                    42             14.0       23.4              255                1.0
                    Ogyrides limicola (D)                       6             2.0       2.0               36                 2.0
                    Nereis succinea (P)                         2             0.7       1.2               12                 3.0
                    Sclerodactyla briareus (HO)                 1             0.3       0.6                 6                5.5
                    Glycera americans (P)                       1             0.3       0.6                 6                5.5
                    Heteromastus filiformis (P)                 1             0.3       0.6                 6                5.5
                    Diopatra cuprea (P)                         1             0.3       0.6                 6                5.5

                    -----------------------------------   Collection Date    30 January 1978  ---------------------------------

                    Mulinia lateralis (M)                       9             3.0       1.7               55                 1.0
                    Heteromastus filiformis (P)                 3             1.0       1.0               18                 2.5
                    Lumbrineris tenuis (P)                      3             1.0       1.0               18                 2.5
                    Nucula proxima. (M)                         2             0.7       1.2               12                 4.0
                    Edotea montosa (1)                          1             0.3       0.6                 6                8.0
                    Pelecypoda (undet.) (M)                     1             0.3       0.6                 6                8.0
                    Nereis succinea (P)                         1             0.3       0.6                 6                8.0
                    Lepidonotus sublevis (P)                    1             0.3       0.6                 6                8.0
                    Paraprionospio pinnata (P)                  1             0.3       0.6                 6                8.0
                    Sabellidae (undet.) (P)                     1             0.3       0.6                 6                8.0
                    Sabella microphthalma (P)                   1             0.3       0.6                 6                8.0

                    ------------------------------------  Collection Date     24 April  1978 ----------------------------------

                    Ampelisca vadorum (A)                     42             14.0       4.4               255                1.0
                    Lumbrineris tenuis (P)                    13              4.3       1.5               79                 2.0
                    Haploscoloplos fragilis (P)               12              4.0       3.0               73                 3.0
                    Leucon americanus (C)                       4             1.3       1.5               24                 4.0
                    Glycinde solitaria (P)                      3             1.0       1.0               18                 5.0
                    Ogyrides limicola (D)                       2             0.7       0.6               12                 7.0
                    Oxyurostylis smithi (C)                     2             0.7       0.6               12                 7.0
                    Paraprionospio pinnata   (P)                2             0.7       0.6               12                 7.0
                    Mulinia lateralis (M)                       1             0.3       0.6                 6               10.5
                    Turridae (undet.) (M)                       1             0.3       0.6                 6               10.5
                    Glycera americana (P)                       1             0.3       0.6                 6               10.5
                    Drilonereis magna (P)                       1             0.3       0.6                 6               10.5

                    ------------------------------------  Collection Date     17 July   1978 -----------------------------------

                    Glycinde solitaria (P)                      8             2.7       1.5               48                1.0
                    Haploscoloplos fragilis (P)                 7             2.3       3.2               42                2.0
                    Paraprionospio pinnata (P)                  3             1.0       0.0               18                3.0
                    Ogyrides limicola (D)                       1             0.3       0.6                 6               6.0
                    Callianassa atlantica (D)                   1             0.3       0.6                 6               6.0
                    Glycera ame'ricana (P)                      1             0.3       0.6                 6               6.0
                    Sigambra tentaculata (P)                    1             0.3       0.6                 6               6.0
                    Lumbrineris tenuis (P)                      1             0.3       0.6                 6               6.0

                                                                         44

                 Table 28. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB14. (A
                            Amphipoda; C = Cumacea; D = Decapoda; I = Isopoda; M = Mollusca: P = Polychaeta).

                                                                         SB14
                                                                                                                                Rank
                                                               Total            Number/0.05m2             Estimated              by
                              Species                         Number             x         SD            Number/m2              Number

                 ------------------------------------    Collection   Date    23 August 1977  ------------------------------------

                 Paraprionospio pinnata (P)                      52             17.3       17.9              315                 1.0
                 Tumbrineris tenuis (P)                          10              3.3       4.9                61                 2.0
                 Glycinde solitaria (P)                          4               1.3       2.3                24                 3.0
                 Periclimenes longicaudatus (D)                  3               1.0       1.7                18                 4.5
                 Rereis succinea (P)                             3               1.0       1.7                18                 4.5
                 Haploscoloplos fragilis (P)                     2               0.7       1.2                12                 6.0
                 fanopeus herbst@ii -(D)                         1               0.3       0.6                  6                9.0
                 Melita nitida (A)                               1               0.3       0.6                  6                9.0
                 Paraphoxus oculatus (A)                         1               0.3       0.6                  6                9.0
                 Mulinia lateralis (M)                           1               0.3       0.6                  6                9.0
                 Heteromastus filiformis     (P)                 1               0.3       0.6                  6                9.0

                 ------------------------------------    Collection Date       31 January 1978 -----------------------------------

                 Streblospio benedictii (P)                      58             19.0       19.1              352                 1.0
                 Sabellaria vulgaris (P)                         43             14.3       22.2              261                 2.0
                 Lumbrineris tenuis (P)                          21              7.0       2.7               127                 3.0
                 Heteromastus filiformis     (P)                    8            2.7       2.1                48                 4.0
                 Rulinia lateralis (M)                              5            1.7       2.1                30                 5.0
                 Eteone heteropoda (P)                              4            1.3       1.5                24                 6.0
                 Ampelisca vadorum (A)                              3            1.0       0.0                18                 9.0
                 Macoma sp. (M)                                     3            1.0       1.0                18                 9.0
                 Tharyx marioni (P)                                 3            1.0       1.7                18                 9.0
                 Nereis succinea (P)                                3            1.0       1.7                18                 9.0
                 -@araprionospio pinnata (P)                        3            1.0       1.0                18                 9.0
                 Melita nitida @A)                                  2            0.7       0.6                12                12.5
                 Leucon americanus (C)                              2            0.7       1.2                12                12.5
                 Callinectes sapidus (D)                            1            0.3       0.6                  6               17.5
                 Caprella equilibra (A)                             1            0.3       0.6                  6               17.5
                 Corophium sp. (A)                                  1            0.3       0.6                  6               17.5
                 Glycera americana (P)                              1            0.3       0.6                  6               17.5
                 Scoloplos rubra (P)                                1            0.3       0.6                  6               17.5
                 Drilonereis magna (P)                              1            0.3       0.6                  6               17.5
                 Thelepus setosus (P)                               1            0.3       0.6                  6               17.5
                 -�abella microphthalma    (P)                      1            0.3       0.6                  6               17.5

                 -------------------------------------     Collection Date     25 April 1978  ------------------------------------
                 Lumbrineris tenuis (P)                          33             11.0       5.3               200                 1.0
                 Ampelisca vadorum (A)                           14              4.7       1.2                85                 2.5
                 Spiophanes bombyx (P)                           14              4.7       5.7                85                 2.5
                 Cirratulidae (undet.) (P)                       11              3.7       4.7                67                 4.0
                 Paraprionospi pinnata (P)                          7            2.3       1.5                42                 5.5
                 Mediomastus cafi7forniensis    (P)                 7            2.3       0.6                42                 5.5
                 -@treblospio benedicti (P)                         6            2.0       2.7                36                 7.0
                 _KP-reis succinea (P)                              4            1.3       2.3                24                 8.0
                 Rucula proxima (M)                                 3            1.0       1.7                18                 9.5
                 Heteromast us filiformis (P)                       3            1.0       1.0                18                 9.5
                 OLcyurostylis smithi (C)                           2            0.7       1.2                12                11.5
                 Drilonereis magna (P)                              2            0.7       0.6                12                11.5
                 Paguridea (undet.) (D)                             1            0.3       0.6                  6               21.5
                 UUca sp. (D)                                       1            0.3       0.6                  6               21.5
                 Unciola serrata (A)                                1            0.3       0.6                  6               21.5
                 Edotea mo'ntosa (1)                                1            0.3       0.6                  6               21.5
                 Actinaria (undet.)                                 1            0.3       0.6                  6               21.5
                 Nemertina (undet.)                                 1            0.3       0.6                  6               21.5
                 Mercenaria mercenaria (M)                          1            0.3       0.6                  6               21.5
                 Iyonsia hyalina (M)                                1            0.3       0.6                  6               21.5
                 Acteocina canaliculata (M)                         1            0.3       0.6                  6               21.5
                 Tellinidae (undet.) M)                             1            0.3       0.6                  6               21.5
                 Sabellaria vulgaris (P)                            1            0.3       0.6                  6               21.5
                 Pectinaria gouldii (P)                             1            0.3       0.6                  6               21.5
                 Glycinde solitaria (P)                             1            0.3       0.6                  6               21.5
                 Diopatra cuprea (P)                                1            0.3       0.6                  6               21.5
                 Eteone heteropoda (P)                              1            0.3       0.6                  6               21.5
                 Exogone dispar (P)
                 Thelepus setosus (P)                               1            0.3       0.6                  6               21.5
                 Haploscoloplos sp. (P)                             1            0.3       0.6                  6               21.5
                                                                    1            0.3       0.6                  6               21.5

                  Table 28.(Cont.)

                                                                       SB14
                                                                                                                           Rank
                                                             Total            Number/0.05m2          Estimated              by
                              Species                       Number            -:w       SD           Number/m2           Number

                  ------------------------------------  Collection  Date   18 August 1978 ------------------------------------

                  Lumbrineris tenuis (P)                        8             2.7       2.9                48               1.0
                  Heteromastus filiformis (P)                   .4            1.3       0.6                24               2.0
                  Marphysa sanguinea (P)                        2             0.7       1.2                12               3.0
                  Nemertina (undet.)                            1             0.3       0.6                 6               5.0
                  Mercenaria mercenaria   (M)                   1             0.3       0.6                 6               5.0
                  Mulinia lateralis (M)                         1             0.3       0.6                 6               5.0

                                                                         46

                  Table 29. Macroinvertebrate species collected from the Sewee Bay channel station, SB15. (A
                             Amphipoda; D = Decapoda; M = Mollusca; 0 = Ophiuroidea; P = Polychaeta).

                                                                        SB15
                                                                                         2                                 Rank
                                                          Total            Number/0.05m             Estimated              by
                             Species                      Number             -k       SD            Number/m 2           Number

                  -----------------------------------   Collection Date    23 August 1977 ----------------------------------

                  Paraprionospio pinnata (P)                131            43.7       11.8              794                1.0
                  Glycinde solitaria (P)                     10              3.3      2.1                61                2.0
                  Lumbrineris tenuis (P)                     8               2.7      3.8                48                3.0
                  Ogyrides limicola (D)                      1               0.3      0.6                 6                7.0
                  Ampelisca verrilli (A)                     1               0.3      0.6                 6                7.0
                  Anadara ovalis (M)                         1               0.3      0.6                 6                7.0
                  Glycera americana   (P)                    1               0.3      0.6                 6                7.0
                  Notomastus lobatus (P)                     1               0.3      0.6                 6                7.0
                  Scoloplos rubra (P)                        1               0.3      0.6                 6                7.0
                  Diopatra cuprep (P)                        1               0.3      0.6                 6                7.0

                  -----------------------------------   Collection Date    30 January 1978  ---------------------------------

                  Paraprionospio pinnata (P)                 9               3.0      3.6                55                1.0
                  Glycera americana (P)                      3               1.0      1.7                18                2.5
                  Scoloplos rubra (P)                        3               1.0      1.0                18                2.5
                  Streblospio benedicti    (P)               2               0.7      1.2                12                4.0
                  Melita sp. (A)                             1               0.3      0.6                 6                7.5
                  Amphiuridae (undet.)   (0)                 1               0.3      0.6                 6                7.5
                  Mulinia lateralis (M)                      1               0.3      0.6                 6                7.5
                  Heteromastus filiformis    (P)             1               0.3      0.6                 6                7.5
                  Oligochaeta (undet.)                       1               0.3      0.6                 6                7.5
                  Aricidea fragilis (P)                      1               0.3      0.6                 6                7.5

                  ------------------------------------   Collection Date     28 April 1978 ----------------------------------

                  Pelecypoda (undet.) (M)                    4               1.3      2.3                24                1.0
                  Tellinidae (undet.) (M)                    2               0.7      1.2                12                3.0
                  Haploscoloplos fragilis    (P)             2               0.7      1.2                12                3.0
                  Clymenella torquata (P)                    2               0.7      0.6                12                3.0
                  Ampelisca vadorum (A)                      1               0.3      0.6                 6                8.5
                  Erichthonius brasiliensis (A)              1               0.3      0.6                 6                8.5
                  Glycera americana (P)                      1               0.3      0.6                 6                8.5
                  Streblospio benedicti (P)                  1               0.3      0.6                 6                8.5
                  Spiophanes bombyx (P)                      1               0.3      0.6                 6                8.5
                  Heteromastus filiformis (P)                1               0.3      0.6                 6                8.5
                  Nereis succinea (P)                        1               0.3      0.6                 6                8.5
                  Paraprionospio pinnata (P)                 1               0.3      0.6                 6                8.5

                  ------------------------------------   Collection Date     31 July 1978 -----------------------------------

                  Paraprionospio pinnata (P)                 18              6.0      2.7               109                1.0
                  Clymenella torquata (P)                    3               1.0      1.7                18                2.0
                  Turbonilla sp. (M)                         2               0.7      1.2                12                3.0
                  Pinnixa sayana (D)                         1               0.3      0.6                 6                6.0
                  Acteocina canaliculata (M)                 1               0.3      0.6                 6                6.0
                  Tellinidae (undet.) (M)                    1               0.3      0.6                 6                6.0
                  Glycera americana (P)                      1               0.3      0.6                 6                6.0
                  Glycinde solitaria (P)                     1               0.3      0.6                 6                6.0

                                                                      47

                  Table 30. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB16. (D
                             Decapoda; I = Isopoda; M = Mollusca; P = Polychaeta),

                                                                         SB16
                                                                                                                                   Rank
                                                                Total            Number/0.05m2              Estimated               by
                                Species                        Number              -:i        SD           Number/m2              Number

                  -----------------------------------    Collection Date       30 August 1977 -------------------------------------

                  Lumbrineris tenuis (P)                         36              12.0         7.6              218                  1.0
                  Haploscoloplos fragilis (P)                    13                4.3        4.2               79                  2.0
                  Nereis succinea (P)                              5               1.7        0.6               30                  3.0
                  -Glycera americana (P)                           4               1.3        2.3               24                  4.0
                  Aricidea fragilis (P)                            3               1.0        1.7               18                  5.0
                  c@emma geuma -(M)                                2               0.7        1.2               12                  6.0
                  Cyathura-'@urbanki (1)                           1               0.3        0.6                 6                10.0
                  -Rercenaria mercenaria   (M)                     1               0.3        0.6                 6                10.0
                  Tagelus divisus (M)                              1               0.3        0.6                 6                10.0
                  Rulinia lateralis (M)                            1               0.3        0.6                 6                10.0
                  Heteroma-stusfiliformis (P)                      1               0.3        0.6                 6                10.0
                  75rilonereis magna (P)                           1               0.3        0.6                 6                10.0
                  Maldanidae (-undet.) (P)                         1               0.3        0.6                 6                10.0

                  ----------------------------------     Collection Date     10 February 1978  ------------------------------------

                  Laeonereis culveri (P)                         82              27.3         13.6             497                  1.0
                  Nereis succinea (P)                            44              14.7         11.0             267                  2.0
                  Heteromastus filiformis (P)                    25                8.3        5.5              152                  3.0
                  Lumbrineris tenuis (P)                         21                7.0        2.0              127                  4.0
                  Glycera americana (P)                            6               2.0        1.0               36                  5.5
                  Eteone heteropoda (P)                            6               2.0        2.7               36                  5.5
                  Haploscoloplos sp. (P)                           5               1.7        2.1               30                  7.0
                  Macoma extenuata (M)                             4               1.3        2.3               24                  9.0
                  Paraprionospio pinnata     (P)                   4               1.3        1.5               24                  9.0
                  Cirratulidae (undet.) (P)                        4               1.3        2.3               24                  9.0
                  Mercenaria mercenaria (M)                        2               0.7        0.6               12                 12.0
                  Mulinia lateralis (M)                            2               0.7        0.6               12                 12.0
                  -@treblospio benedicti   (P)                     2               0.7        1.2               12                 12.0
                  Gemma gemma (M)                                  1               0.3        0.6                 6                14.5
                  Pectinaria gouldii (P)                           1               0.31       0.6                 6                14.5

                  ------------------------------------    Collection Date       25 April 1978 -------------------------------------

                  Lumbrineris tenuis (P)                         18                6.0        4.6              109                  1.0
                  Heteromastus filiformis (P)                      8               2.7        1.5               48                  2.5
                  Mediomastus californiensis    (P)                8               2.7        2.1               48                  2.5
                  Streblospio benedicti (P)                        7               2.3        2.5               42                  4.5
                  Nereis succinea (P)                              7               2.3        4.0               42                  4.5
                  Uaploscoloplos-fragilis (P)                      6               2.0        2.0               36                  6.5
                  Paraprionospio pinnata (P)                       6               2.0        1.0               36                  6.5
                  Tellinidae (und-et.)(M)                          4               1.3        1.5               24                  8.5
                  Cirratulidae (undet.) (P)                        4               1.3        2.3               24                  8.5
                  Nemertina (undet.)                               2               0.7        0.6               12                 11.0
                  Mercenaria mercenaria (M)                        2               0.7        0.6               12                 11.0
                  dlycera americana (P)                            2               0.7        0.6               12                 11.0
                  Pinnixa sayana (D)                               1               0.3        0.6                 6                16.5
                  Gemma gemma (M)                                  1               0.3        0.6                 6                16.5
                  Mulinia lateralis (M)                            1               0.3        0.6                 6                16.5
                  Acteocina canal-i-culata (M)                     1               0.3        0.6                 6                16.5
                  Scoloplos rubra (P)                              1               0.3        0.6                 6                16.5
                  Glycinde solitaria (P)                           1               0.3        0.6                 6                16.5
                  Drilonereis magna (P)                            1               0.3        0.6                 6                16.5
                  Notomastus sp. (P)                               1               0.3        0.6                 6                16.5

                  -----------------------------------    Collection Date      18 August 1978  -------------------------------------

                  Heteromastus filiformis (P)                    38              12.7         21.9             230                  1.0
                  Uca pugilator (D)                              12                4.0         5.3              73                  2.0
                  Lumbrineris tenuis (P)                           6               2.0         3.5              36                  3.0
                  Gemma gemma (M)                                  5               1.7         1.5              30                  4.0
                  Nereis succinea (P)                              4               1.3         1.5              24                  5.0
                  Cirratulidae (undet.) (P)                        2               0.7         1.2              12                  6.0
                  Palaemonetes sp. (D)                             1               0.3         0.6                6                 7.0

                                                                               48

                      Table 31. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB17. (A
                                  Amphipoda; C = Cumacea; D = Decapoda; Ho = Holothoroidea; I = Isopoda; M                 Mollusca;
                                  0 = Ophiuroidea; P = Polychaeta).

                                                                            SB17
                                                                                                                                      Rank
                                                                 Total              Number/0.05m2             Estimate                 by
                                  Species                       Number               _9        SD             Number/m                Number

                    ----------------------------------      Collection Date      30 August 1977    ------------------------------------

                    Lumbrineris tenuis (P)                         49               16.3       4.5                 297                 1.0
                    Caprella equilibra (A)                           4               1.3       2.3                 24                  2.0
                    Glycera americana (P)                            3               1.0       1.0                 18                  3.5
                    Nereis succinea (P)                              3               1.0       1.0                 18                  3.5
                    Hemipholis elongata (0)                          2               0.7       1.1                 12                  7.0
                    Macoma sp. (M)                                   2               0.7       1.1                 12                  7.0
                    Scoloplos rubra (P)                              2               0.7       1.1                 12                  7.0
                    Arabella iricolor (P)                            2               0.7       1.1                 12                  7.0
                    Aricidea fragilis (P)                            2               0.7       0.6                 12                  7.0
                    Upogebia affinis (D)                             1               0.3       0.6                   6                13.5
                    Pagurus pollicaris (D)                           1               0.3       0.6                   6                13.5
                    Xanthidae (undet.) (D)                           1               0.3       0.6                   6                13.5
                    Sclerodactyla briareus     (Ho)                  1               0.3       0.6                   6                13.5
                    Nucula proxima (M)                               1               0.3       0.6                   6                13.5
                    Eunice websteri (P)                              1               0.3       0.6                   6                13.5
                    Diopatra cuprea (P)                              1               0.3       0.6                   6                13.5
                    Drilonereis magna (P)                            1               0.3       0.6                   6                13.5

                    ---------------------------------      Collection Date      10 February 1978   -----------------------------------

                    Haploscoloplos sp. (P)                          11               3.7       1.5                 67                  1.0
                    Ogyrides limicola (D)                            4               1.3       2.3                 24                  2.5
                    Ilyanassa obsoleta (M)                           4               1.3       2.3                 24                  2.5
                    Lumbrineris tenuis (P)                           3               1.0       1.0                 18                  5.0
                    Nereis succinea (P)                              3               1.0       1.0                 18                  5.0
                    Paraprionospio pinnata     (P)                   3               1.0       1.0                 18                  5.0
                    Paraphoxus spinosus (A)                          2               0.7       0.6                 12                  9.5
                    Ampelisca abdita (A)                             2               0.7       1.2                 12                  9.5
                    Acteocina canaliculata     (M)                   2               0.7       1.2                 12                  9.5
                    Macoma extenuata (M)                             2               0.7       1.2                 12                  9.5
                    Glycera americana (P)                            2               0.7       0.6                 12                  9.5
                    Glycinde solitaria (P)                           2               0.7       0.6                 12                  9.5
                    Corophium acherusicum     (A)                    1               0.3       0.6                   6                16.0
                    Caprella equilibra (A)                           1               0.3       0.6                   6                16.0
                    Leucon americanus (C)                            1               0.3       0.6                   6                16.0
                    Edotea montosa (1)                               1               0.3       0.6                   6                16.0
                    Macoma sp. (M)                                   1               0.3       0.6                   6                16.0
                    Laeonereis culveri (P)                           1               0.3       0.6                   6                16.0
                    Eteone heteropoda (P)                            1               0.3       0.6                   6                16.0

                    -----------------------------------      Collection Date      25 April 1978    ------------------------------------

                    Haploscoloplos fragilis (P)                     17               6.0       5.0                 103                 1.0
                    Ampelisca vadorum (A)                            9               3.0       1.0                 55                  2.0
                    Acteocina canaliculata (M)                       5               1.7       1.2                 30                  3.5
                    Lumbrineris tenuis (PT                           5               1.7       1.5                 30                  3.5
                    Tellinidae (undet.) (M)                          4               1.3       1.5                 24                  5.0
                    Paraprionospio pinnata     (P)                   3               1.0       1.0                 18                  6.0
                    Glycera americana (P)                            2               0.7       0.6                 12                  7.0
                    Ogyrides limicola (D)                            1               0.3       0.6                   6                11.0
                    Leucon americanus (C)                            1               0.3       0.6                   6                11.0
                    Heteromastus filiformis (P)                      1               0.3       0.6                   6                11.0
                    Glycinde solitaria (P)                           1               0.3       0.6                   6                11.0
                    Drilonereis magna (P)                            1               0.3       0.6                   6                11.0
                    Orbiniidae (undet.) (P)                          1               0.3       0.6                   6                11.0
                    Notomastus sp. (P)                               1               0.3       0.6                   6                11.0

                    ----------------------------------      Collection Date      17 August 1978    ------------------------------------

                    Glycera americana (P)                            2               0.7       0.6                 12                  1.0
                    Lumbrineris tenuis (P)                           1               0.3       0.6                   6                 2.0

                                                                            49

                        Table 32. Macroinvertebrate species collected from the Sewee Bay subtidal station, SB18. A
                                   Amphipoda; C = Cumacea; D = Decapoda; I = Isopoda; M = Mollusca; My = Mysidacea;
                                   0 = Ophiuroidea; P = Polychaeta; S = Stomatopoda).

                                                                        SB18
                                                                                            2                                Rank
                                                              Total             Number/0.05m            Estimated             by
                              Species                        Number             -9       SD            Number/m2             Number

                  ------------------------------------   Collection  Date   25 August 1977  ------------------------------------

                  Lumbrineris tenuis (P)                       47               15.7     16.1              285                1.0
                  Glycinde solitaria (P)                         3              1.0      0.0                18                2.0
                  Squilla empusa (S)                             1              0.3      0.6                  6               6.5
                  Ampel,isca vadorum (A)                         1              0.3      0.6                  6               6.5
                  Cyathura burbanki    (1)                       1              0.3      0.6                  6               6.5
                  Scoloplos rubra (P)                            1              0.3      0.6                  6               6.5
                  Heteromastus filiformis (P)                    1              0.3      0.6                  6               6.5
                  Drilonereis magna (P)                          1              0.3      0.6                  6               6.5
                  Aricidea fragilis (P)                          1              0.3      0.6                  6               6.5
                  Syllidae (undet.) (P)                          1              0.3      0.6                  6               6.5

                  ------------------------------------   Collection Date    31  January 1978 -----------------------------------

                  Mulinia lateralis (M)                        28               9.3      3.1               170                1.0
                  Paraprionospio pinnata (P)                   22               7.3      4.7               133                2.0
                  Ogyrides limicola (D)                        21               7.0      7.6               127                3.0
                  Paracaprella tenuis (A)                      18               6.0      10.4              109                4.0
                  Erichthonius brasiliensis (A)                13               4.3      7.5                79                5.0
                  Tharyx marioni (P)                             6              2.0      3.5                36                6.5
                  Heteromastus filiformis (P)                    6              2.0      2.0                36                6.5
                  Lumbrineris tenuis (P)                         5              1.7      2.1                30                8.0
                  Ampelisca vadorum (A)                          4              1.3      1.5                24               10.0
                  Batea catharinensis (A)                        4              1.3      2.3                24               10.0
                  Nereis succinea (P)                            4              1.3      1.5                24               10.0
                  Microdeutopus gryllotalpa (A)                  3              1.0      1.7                18               12.5
                  Oligochaeta (undet.)                           3              1.0      1.0                18               12.5
                  Caprella equilibra (A)                         2              0.7      1.2                12               15.0
                  Melita sp. (A)                                 2              0.7      1.2                12               15.0
                  Haploscoloplos robustos (P)                    2              0.7      0.6                12               15.0
                  Pinnixa sp. (D)                                1              0.3      0.6                  6              22.5
                  Corophium acherusicum (A)                      1              0.3      0.6                  6              22.5
                  Leucon americanus (C)                          1              0.3      0.6                  6              22.5
                  Cyathura burbanki (1)                          1              0.3      0.6                  6              22.5
                  Abra aequalis (M)                              1              0.3      0.6                  6              22.5
                  Acteocina canaliculata (M)                     1              0.3      0.6                  6              22.5
                  Pectinaria gouldii (P)                         1              0.3      0.6                  6              22.5
                  Eteone heteropoda (P)                          1              0.3      0.6                  6              22.5
                  Odontosyllis fulgurans (P)                     1              0.3      0.6                  6              22.5
                  Aricidea fragilis (P)                          1              0.3      0.6                  6              22.5
                  Ampharetidae (undet.) (P)                      1              0.3      0.6                  6              22.5
                  Sabella microphthalma (P)                      1              0.3      0.6                  6              22.5

                  -------------------------------------   Collection Date    24 April 1978  ------------------------------------

                  Ampelisca vadorum (A)                        23               7.7      1.5               139                1.0
                  Paraprionospio pinnata (P)                   11               3.7      2.1                67                2.0
                  Lumbrineris tenuis (P)                         7              2.3      1.5                42                3.0
                  Ogyrides limicola (D)                          6              2.0      1.0                36                4.5
                  Spionidae (undet.) (P)                         6              2.0      2.0                36                4.5
                  Neomysis americana (MY)                        5              1.7      2.1                30                6.5
                  Tellinidae (undet.) (M)                        5              1.7      1.5                30                6.5
                  Glycera americana (P)                          4              1.3      1.2                24                9.0
                  Glycinde solitaria (P)                         4              1.3      0.6                24                9.0
                  Haploscoloplos sp. (P)                         4              1.3      0.6                24                9.0
                  Batea cathariensis (A)                         2              0.7      1.2                12               14.0
                  Paracaprella tenuis (P)                        2              0.7      1.2                12               14.0
                  Leucon americanus (C)                          2              0.7      0.6                12               14.0
                  Oxyurostylis smithi   (C)                      2              0.7      0.6                12               14.0
                  Gammaridae (undet.)   (A)                      2              0.7      0.6                12               14.0
                  Hemipholis elongata   (0)                      2              0.7      0.6                12               14.0
                              Cu-nd                                             0.7      0.6                12
                  Polynoidae      et.) (P)                       2                                                           14.0
                  Edotea montosa (P)                             1              0.3      0.6                12               20.5
                  Nucula proxima (M)                             1              0.3      0.6                12               20.5
                  Lyonsi hyalina (P)                             1              0.3      0.6                12               20.5
                  Sabellaria vulgaris (P)                        1              0.3      0.6                12               20.5
                  Oligochaeta Fun-det.)                          1              0.3      0.6                12               20.5
                  Cirratulidae (undet.) (P)                      1              0.3      0.6                12               20.5

                 Table 32.(Cont.)

                                                                           SB18

                                                                                                                                   Rank
                                                               Total              Number/0.05m2             Estimatel               by
                               Species                         Number              -x        SD             Number/m             Number

                 ------------------------------------     Collection   Date    17  August 1978 ------------------------------------

                 Lumbrineris tenuis (P)                           7                2.3       0.6               42                   1.0
                 Panopeus herbstii (D)                            4                1.3       0.6               24                   2.5
                 -Tharyx marioni (P)                              4                1.3       1.2               24                   2.5
                 Relita appendiculata     (A)                     3                1.0       1.7               18                   4.0
                 Cyathura burbanki (1)                            2                0.7       0.6               12                   6.5
                 Scoloplos sp. (P)                                2                0.7       1.2               12                   6.5
                 Glycinde solitaria (P)                           2                0.7       0.6               12                   6.5
                 Paraprionospio pinnata     (P)                   2                0.7       0.6               12                   6.5
                 'Remertina (undet-.)                             1                0.3       0.6                6                  13.5
                 Hemipholis elongata (0)                          1                0.3       0.6                6                  13.5
                 Nucula proxima (M)                               1                0.3       0.6                6                  13.5
                 �abella-r-iaZ,j@ilaris (P)                       1                0.3       0.6                6                  13.5
                 Pista cristata (F)                               1                0.3       0.6                6                  13.5
                 Heteromastus filiformis     (P)                  1                0.3       0.6                6                  13.5
                 Nereis succinea (P)                              1                0.3       0.6                6                  13.5
                 -Krabelf--
                         a tricolor (P)                           1                0.3       0.6                6                  13.5
                 Notomastus sp. (P)                               1                0.3       0.6                6                  13.5
                 Polynoidae (undet.) (P)                          1                0.3       0.6                6                  13.5

               Table 33. Species diversity, evenness and richness values for Sewee Bay grab samDles from August 1977 to August 1978.             First Quarter = samples
                          collected before dredging; Second Quarter = samples collected immediately after dredging; Third Quarter.= samples collected 3 months
                          after dredging; Fourth Quarter      samples collected 6 months after dredging and 1 year after first quarter samples.

                                         Diversity (H')                                   Evenness (J')                                   Richness (SR)
                            First       Second      Third       Fourth      First       Second      Third       Four@h      First        Second      Third      Tourth
               Station     Quarter      Quarter    Quarter      Quarter    Quarter     Quarter     Quarter      Quarter    Quarter     Quarter      Quarter     Quarter     Station
                SBOl        2.06        2.02        3.61        1.63        0.65        0.72        0.76        0.58        1.91         1.73        5.99        1.76        SBOl
                SB02        1.95        2.73        0.18        3.03        0.69        0.86        0.11        0.96        1.63         2.35        0.44        2.89        SB02
                SB03        0.44        3.66        3.85        3.48        0.28        0.94        0.85        0.94        0.60         4.25        4.99        3.80        SB03
                SB04        3.24        2.7.3       3.03        1.97        0.90        0.74        0.80        0.62        3.20         3.35        3.42        1.95        SB04
                SB05        2.19        3.29        2.98        2.48        0.63        0.89        0.86        0.96        2.63         3.83        2.79        2.09        SB05
                SB06        2.35        3.02        2.23        2.32        M8          0.91        0.70        1.00        2.08         2.83        1.94        2.48        SB06

                SB07        3.30        4.29        3.84        3.65        0.89        0.90        0.84        0.86        3.68         6.65        5.45        4.56        SB07

                SB08        2.76        3.78        3.08        1.57        0.87        0.91        0.74        0.61        2.55         4.17        3.81        1.28        SB08

                SB09        2.45        2.91        2.91        2.07        0.74        0.61        0.75        0.74        1.90         5.24        2.85        1.94        SB09

                SB10        2.48        1.81        3.65        2.27        0.67        0.78        0.77        0.63        2.72         1.52        4.94        2.24        SB10

                SBll        1.89        2.09        2.70        2.16        0.60        0.61        0.75        0.72        0.47         2.09        2.32        1.48        SBll

                            1.78        2.31        3.07        2.11        0.69        0.73        0.77        0.70        1.17         1.83        3.46        1.84        SB12

                SB13        1.24        2.92        2.38        2.42        0.44        0.84        0.67        0.81        1.50         3.14        2.48        2.23        SB13

                SB14        1.88        2.94        3.83        2.09        0.54        0.67        0.78        0.81        2.29         3.91        6.01        1.76        SB14

                SB15        1.01        2.78        3.39        1.89        0.30        0.84        0.95        0.63        1.78         2.87        3.81        2.10        SB15

                SB16        2.41        2.72        3.73        1.97        0.65        0.70        0.87        0.70        2.82         2.62        4.31        1.42        SB16

                SB17        2.34        3.85        3.08        0.92        0.57        0.91        0.81        0.92        3.68         4.68        3.29        0.91        SB17

                SB18        1.27        3.86        3.90        3.83        0.38        0.81        0.86        0.92        2.22         5.35        4.83        4.74        SB18

        Table 34. Mean abundance and total number of species collected in the Sewee Bay grab samples from August 1977 to
                   August 1978. Mean values indicate number/0.05m2. First Quarter = samples collected before dredging;
                   Second Quarter = samples collected immediately after dredging; Third Quarter = samples collected 3
                   months after dredging; Fourth Quarter = samples collected 6 months after dredging and 1 year after
                   first quarter samples.

                         FIRST QUARTER                 SECOND QUARTER                  THIRD QUARTER                FOURTH QUARTER
        Station    3F Abundance   No. Species    iF Abundance    No. Species    -x Abundance    No. Species    3F Abundance   No. Species

         SBOl         22.0               9           10.7               7           25.7             27           10.0                7

         SB02         13.3               7           10.0               9           30.0               3            5.3               9

         SB03           9.3              3            9.0              15           27.3             23             8.0             13

         SB04         10.3              12           12.0              13           15.0             14           20.3                9

         SB05         15.0              11            7.7              13           12.0             11             3.7               6

         SB06           9.7              8            8.0              10           20.7               9            1.7               5

         SB07         17.3              13           16.7              27           22.7             24           17.3              19

         SBO8           7.7              9           19.7              18           29.0             18           16.3                6

         SB09         38.0              10           47.7              27           45.7             15             7.3               7

         SB10         27.3              13            4.7               5           64.3             29           45.7              12

         SBll         53.7               9           39.3              11           37.7             12           38.3                8

         SB12         23.7               6           26.7               9           25.3             16           15.0                8

         SB13         18.0               7            8.0              11           28.0             12             7.7               8

         SB14         26.3              11           55.3              21           41.3             30             5.7               6

         SB15         52.0              10            7.7              10             6.0            12             9.3               8

         SB16         23.3              13           69.7              15           27.3             20           22.7                7

         SB17         25.7              17           15.7              19           17.3             14             1.0               2

         SB18         19.3              10           51.7              28           31.7             23           12.0              18

                                                                       53

           Table 35. Wet-weight biomass of Sewee Bay grab samples from August 1977 to August 1978. Values represent mean
                      and standard deviation of biomass for all species collected/0.05m2, First Quarter = samples collected before
                      dredging; Second Quarter = samples collected immediately after dredging; Third Quarter = samples
                      collected 3 months after dredging; Fourth Quarter       samples collected 6 months after dredging and 1
                      year after first quarter samples.

                                    FIRST                       SECOND                       THIRD                      FOURTH
                                    QUARTER                     QUARTER                      QUARTER                     QUARTER
           Station             3E           SD                          SD             _iE          SD              3E          SD

              SBOl            0.26         0.17           0.16         0.15           0.86          0.94           0.09         0.06

              SB02            0.06         0.02           0.13         0.64           0.72          0.53           0.06         0.08

              SB03            0.06         0.08           0.13         0.09           0.16          0.11           0.41         0.16

              SB04            0.50         0.55           0.34         0.45           0.22          0.11           0.10         0.03

              SB05            0.81         0.56           1.46         2.29           0.11          0.08           0.27         0.39

              SB06            0.40         0.55           1.37         1.55           0.18          0.02           0.07         0.11
              SB07            0.14         0.13           2.24         3.78           1.78          2.74           0.56         0.31
              SB08            0.12         0.14           0.20         0.07           0.16          0.05           0.22         0.16
              SB09            0.40         0.35           0.30         0.28           0.19          0.03           0.07         0.06
              SB10            0.25         0.22           0.04         0.02           0.33          0.08           1.41         0.91
              SBll            0.29         0.72           1.53         2.32           0.70          0.67           0.38         0.21
              SB12            0.47         0.23           0.23         0.09           0.98          1.48           0.14         0.09
              SB13            5.25         8.82           0.07         0.06           0.20          0.04           0.09         0.11
              SB14            2.89         4.96           0.49         0.35           0.27          0.26           0.32         0.55
              SB15            7.02         11.59          0.26         0.27           0.09          0.12           0.49         0.16
              SB16            0.25         0.29           0.60         0.67           0.34          0.17           1.67         2.17
              SB17            2.34         2.63           2.38         3.99           0.35          0.28           0.05         0.04
              SB18            0.08         0.06           0.28         0.13           0.74          0.10           0.54         0.50

                                                                            54

                                            Immill Illm
                                             3 6668 14104 1212